Muisca heppneri, Opitz, Pitz Pinola Weston, 2018

Opitz, Pitz Pinola Weston, 2018, Classification, Natural History, and Evolution of the Subfamily Peloniinae O (Coleoptera: Cleroidea: Cleridae). Part IX. Taxonomic revision of the New World genus Muisca S, Linzer biologische Beiträge 50 (1), pp. 587-653 : 618

Muisca heppneri OPITZ nov.sp. ( Figs 30 , 60, 95, 112, 142)

Holotype: ♀. PANAMÁ: Pmá. Prov. , Cerro Campana, 820 m, 8°40'N, 79°56'W. A second label reads: 5 Sept. ’77, H. P. Stockwell ( FSCA) . Paratypes: 19 specimens. Costa Rica: Provincia de Guanacaste, 9 km S Santa Ceciliam Parque Nacional Guanacaste, 24-VIII-1992, P Rios ( WOPC, 1). Panamá: Provincia de Veragua , Santa Fe , Alto Piedra , 11-16-VI- 2010, 850 m, J. B. Heppner ( FSCA, 1) ; Provincia de Panamá, Cerro Campana , 8°40'N, 79°56'W, 850 m, 2-IX-1972, H. Stockwell ( WOPC, 2) ; idem, 5-IX-1977, H. P Stockwell ( WOPC, 1) ; idem, 18-VII-1970, H. P. Stockwell ( WOPC, 1) ; idem, 13-V-1978, 2700', C. W. & L. B. O’Brien & Marshall ( WOPC, 5) ; idem, 18-VI-1976, W. E. Clark ( WOPC, 5) ; idem, 16-VII-1976, W. Clark ( WOPC, 1) ; idem, 6-V- 1973, 454 m, O’Brien & Marshall ( WOPC,1) ; idem, 29-VI- 1974, L. & C. O’Brien, Marshall, & H. Stockwell ( WOPC, 1) .

D i a g n o s i s: Muisca heppneri specimens may be distinguished from those of its sister species, M. lateripunctata , by coloration near the elytral humeral angle. In M. heppneri specimens there are 2 distinct black maculae near the humeral angle. In M. lateripunctata specimens the humeral region is marked by 1 irregularly-shaped brown macula.

D e s c r i p t i o n: Size: Length 6.0 mm; width 2.5 mm. Form: As in Fig. 142 . Color: Light castaneous, except each elytron with 2 black spots near the humeral angle and a black angular fascia at middle of disc. Head: Funicular antennomeres filiform, capitulum about as long as combined length of funicular antennomeres, capitular antennomeres 9 and 10 oblong triangular ( Fig. 30 ), antennomere 11 obovate; frons wider than width of eye ( EW /FW 30/35). Thorax: Pronotum (Fig. 60) oblong ( PW /PL 95/115), side margin with well-developed tubercle, disc subglabrous at middle; elytral asetiferous punctation striate to distal 2/3 rd ( EL /EW 280/90). Abdomen: Pygidium subquadrate, posterior margin slightly concave; distal margin of 6 th visible sternite broadly concave; aedeagus phallobase wide in basal half, tapered to lobe apices, phallic apex ( Fig. 112 ) triangular.

V a r i a t i o n: Size: Length 5.0- 8.5 mm; width 1.5-3.0 mm. Other than body size the available beetles are quite similar in appearance.

N a t u r a l h i s t o r y: Specimens were collected during May through September at altitudes ranging from 800 to 850 m; most were taken with a beating sheet.

E t y m o l o g y: The specific epithet, heppneri, is a dedicative patronymic. It honors John B. Heppner for his many contributions to taxonomic entomology, among which is his dedication to field taxonomy.

D i s t r i b u t i o n ( Fig.95 ): This species is known only from Panamá.

Figures

Figs 95: Geographic distribution of Muisca species as listed.

Figs 14-33: Antennae. (14) Muisca dilatata, male. (15) M. apicalis, male. (16) M. hirtula, male. (17) M. xanthura, male. (18) M. angulicollis, male. (19) M. signa, male. (20) M. fera, male. (21) M. octonotata, male. (22) M. maculosa, male. (23) M. anachyma, male. (24) M. insigna, male. (25) M. dozieri, male. (26) M. togata, male. (27) M. irrorata, male. (28) M. menda, female. (29) M. hexa, female. (30) M. heppneri, male. (31) M. magdalena, female. (32) M. mestolinea, male. (33) M. omma, male.

Figs 141-144: Habitus. (141) Muisca omma. (142) M. heppneri. (143) M. lateripunctata. (144) M. magdalena.

Figs 107-115: Phalli. (107) Muisca signa. (108) M. angulicollis. (109) M. maculosa. (110) M. mestolinea. (111) M. omma. (112) M. heppneri. (113) M. lateripunctata. (114) M. octonotata. (115) M. tetraspilota.