Tapinopterus (Molopsis) machardi ( Jeanne, 2005 )

Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard & Lassalle, Bernard, 2012, Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species, Zootaxa 3185, pp. 1-35 : 11-13

Tapinopterus (Molopsis) machardi ( Jeanne, 2005)

(Figs. 13, 35, 53, 63, 85, 100, 115, 129)

Type locality. Turquie, Antalya, Col Karaovabeli, 1560 m.

Taxonomic decisions. Molopsis machardi Jeanne, 2005: 383 .

Tapinopterus (Molopsis) machardi ( Jeanne, 2005) , combinatio novum of Molopsis machardi Jeanne, 2005 .

References. Molopsis machardi Jeanne, 2005: 383 , 385.

Material examined. Four specimens.

Type material. One paratype 3 of Tapinopterus (Molopsis) machardi, Col. KARAOVABELI 1400 Antalya Turquie, 0 3.05.96 B.Lassalle (p)// PARATYPE (red p) / Molopsis machardi m. Jeanne det. 2002 (p) (Fig.13) (cBL).

Other material examined. 3 33, Col. KARAOVABELI 1400 Antalya Turquie, 0 3.05.96 G. Dubault & D.Echaroux (cGD, cDE).

Diagnosis. BL 14.5–16.2 mm.

Male genitalia. Median lobe of aedeagus long in dorsal view, apical lamella long, straight, and thin, apical orifice deflected to left lateral position (Figs. 35); basal bulb relatively long, with concave basal orifice, apical part longer than basal one, curved copulatory sclerite in subapical position, apex becoming thinner towards tip, hardly bent downwards (Figs. 53, 63). Right paramere long, regularly curved medially, thicker basally than apically, with well-differentiated, curved subbasal part, apical part shortly acute towards apex (Figs. 85, 100). Left paramere with short transverse apophysis and long oblique paramedial apophysis (Figs. 115, 129).

Female genitalia. Unknown.

This species differs from all other species of Molopsis by the presence of very fine setae underneath the onychium and by the specific shape of the median lobe of aedeagus and right paramere (Figs. 35, 53, 63, 85, 100).

Taxonomic notes. The single paratype studied possesses different numbers of setae underneath the onychium, respectivelly: 1+1 (left anterior leg), 1+1 (right anterior leg), 1+1 (left middle leg), 1+0 (right midlle leg), 2+1 (left posterior leg), 1+0 (right posterior leg). The setae vary in size and thickness, and generally are shorter and finer than typical setae in the most species of Tapinopterus s.l. We consider that the presence / absence of ventral setae on onychium may have different phyletic value to specify monophyletic lines in Pterostichini . This process reflects morphological regressions appeared in different, unrelated clades of the tribe as a result of an adaptation to different environments. The two states of this character occur either in species from different subgenera of the same genus ( Ancholeus Dejean, 1828 and Poecilus Bonelli, 1810 of Poecilus s.l.) or among species from one and same subgenus— Phonias Gozis, 1886 (of the genus Pterostichus Bonelli, 1810 ; cfr. Hůrka, 1996), Molopsis Schatzmayr (this paper).

Distribution. South Taurus, Karaovabeli pass between Kaş and Elmali, W of Antalya (“Lycian Taurus”).

PLATE 2. Habitus of Tapinopterus species (Figs. 13–24). Fig. 13. Tapinopterus (Molopsis) machardi (paratype); Fig. 14. T. (Molopsis) oyukluensis sp. nov. (holotype); Fig. 15. T. (Molopsis) molopinus (holotype of Feronia molopina ); Fig. 16. T. (Molopsis) molopinus (holotype of Tapinopterus dipojranus brussanus ); Fig. 17. T. (Molopsis) molopinus ( Turkey, Uludaġ); Fig. 18. T. (Molopsis) chaudoiri sp. nov. (paratype, Domanic); Fig. 19. T. (Molopsis) chaudoiri sp. nov. (paratype, Oylat); Fig. 20. T. (Molopsis) chaudoiri sp. nov. (paratype, Gök daġlari); Fig. 21. T. (Molopsis) wiedemanni (lectotype); Fig. 22. T. (Molopsis) wiedemanni ( Turkey, Abant); Fig. 23. T. (Molopsis) wiedemanni ( Turkey, Sundiren daġlari); Fig. 24. T. (Molopsis) relegatus sp. nov. (holotype).

PLATE 3. Median lobe of aedeagus (Figs. 25–36), dorsal view (Figs. 25–28, 30–36) and ventral view (Fig. 29) of Tapinopterus species. Fig. 25. Tapinopterus (Pseudomolopsis) rebellis ( Greece, Kalamata); Fig. 26. T. (Molopsis) molopiformis (Tmolos- Gbg., Lydien, West-Kleinasien, Weirather); Fig. 27. T. (Molopsis) molopiformis (Coll Türk 1888); Figs. 28–29. T. (Molopsis) aenigmaticus sp. nov. (holotype); Fig. 30. T. (Molopsis) phrygius (syntype); Fig. 31. T. (Molopsis) phrygius (syntype of T. phrygius pisidicus ); Fig. 32. T. (Molopsis) phrygius (holotype of T. dipojranus ); Fig. 33. T. (Molopsis) phrygius ( Turkey, Beyşehir); Fig. 34. T. (Molopsis) phrygius (paratype of Tapinopterus dipojranus cilicius ); Fig. 35. T. (Molopsis) machardi (paratype); Fig. 36. T. (Molopsis) oyukluensis sp. nov. (holotype). Scale bar: 1 mm.