Mizotrechus fortunensis, Erwin, 2011

Erwin, Terry L., 2011, Rainforest understory beetles of the Neotropics, Mizotrechus Bates 1872, a generic synopsis with descriptions of new species from Central America and northern South America (Coleoptera, Carabidae, Perigonini), ZooKeys 145, pp. 79-128 : 93-94

Fortuna trough beetle Mizotrechus fortunensis sp. n. Figs 9 31 39

Holotype.

PANAMÁ, Chiriquí Province, Fortuna, 1050 m, 8.7341°N, 82.2701°W, 13 August 1978 (H. Wolda)(NMNH: ADP124968, male).

Derivation of specific epithet.

The epithet “fortunensis” is a Latinized noun in apposition, based on a geographic name formed from “Fortuna“ ( Panamá) the place in which adults of this species have been found, and the Latin suffix “-ensis,“ meaning of, or from that place.

Proposed English vernacular name.

Fortuna trough beetle.

Diagnosis.

With the attributes of the genus as described above and large sized for the genus as it is presently understood; adults have castaneous integument, except anterior parts of mandible, baso-lateral corner of labrum, and clypeal suture piceous. Frons smooth, not or barely rugose near eye, surface punctulate. Occiput smooth, punctulate. Pronotum quadrate with lateral margins basally shallowly emarginated, hind angles slightly toothed; base densely microrugose. Elytra broad and short, much broader than width of pronotum across anterior third, and with 3 well-impressed interneurs, others shallower toward margin; margins behind humeri shallowly serrulate. Foreleg femur serrate on postero-ventral margin.

Description.

( Figs 9 , 31 ). Size: See Appendix 1. Large-sized for genus, ABL = 8.5-8.7 mm, SBL = 7.02-7.66 mm, TW = 2.93 -3.03 mm. Color: see diagnosis, above. Luster: Head, pronotum, and legs shiny; elytra dull due to marked slightly stretched well-impressed microsculpture. Head: Labrum quadrate, entire apically. Eye moderately large, moderately convex. Gena moderately long, straight. Frons, occiput, and gena glabrous. Prothorax: Quadrate, narrowed slightly toward base, margin beaded, not explanate except at hind angle, hind angle feebly dentate; surface punctulate, punctures widespread, and with microsculpture nearly effaced, surface glabrous. Pterothorax: Elytron moderately convex, disk flat, intervals flat, interneurs striate, not punctate, shallower toward margin, apex slightly oblique, straight, sutural apex narrowly and slightly rounded. Metasternum sparsely setiferous in male. Legs: Normal in both sexes; foreleg femur (as in Fig. 24 ) serrate on postero-ventral margin; posterior trochanter narrowly rounded in both sexes, length half that of femur. Abdomen: Abdominal sterna moderately setiferous; sternum IV of male without medial patch of decumbent setae; sternum VII medially and shallowly V-notched in male. Male genitalia: Median lobe ( Fig. 31 ) short and robust with ostium moderately elongate, over half the length of the median lobe; apex with a very short and narrow distal end, slightly rounded in lateral aspect, less twisted laterally than in Mizotrechus costaricensis , ventral margin proximal to apex straight then briefly curved to apex; endophallus with complexly folded tracheal fields; phallobase hooded, opening more or less 20 degrees off axis of shaft. Parameres large, left twice longer than the right, both broadly rounded, asetose. Female genitalia: Unknown.

Dispersal potential.

These beetles are macropterous and capable of flight. They are moderately swift and agile runners.

Way of life.

The six adults were collected at UV light trap in premontane forest understory. Adults are active in May, July, and August, in the rainy and early dry seasons.

Other specimens examined.

Paratypes: Panamá, Chiriqui Province, Fortuna, 1050 m, 8.7341°N, 82.2701°W, 15 May 1978 (H. Wolda)(NMNH: ADP124964, male), 7 July 1978 (H. Wolda)(NMNH: ADP124964, male), 19 July 1978 (H. Wolda)(NMNH: ADP124952, male), 27 July 1977 (H. Wolda)(NMNH: ADP070275, male), 13 August 1978 (H. Wolda)(NMNH: ADP124954, male).

Geographic distribution.

( Fig. 39 ). This species is currently known only from the type locality in the premontane forested midlands of Panamá.

Figures

Plate 3. Digital Photo-illustrations, habitus, dorsal aspect: 9 Mizotrechus fortunensi s sp. n., ABL = 8.7 mm, male holotype, ADP 124968; type locality. 10 Mizotrechus gorgona. sp. n., ABL = 8.4 mm, female Holotype, ADP 128622; type locality. 11 Mizotrechus grossus sp. n., dorsal aspect, ABL = 8.9 mm, male paratype, ADP 127169; type locality. 12 Mizotrechus jefe sp. n., ABL = 6.6 mm, male paratype, ADP 011173; type locality.

Plate 6. Digital Photo-illustrations, anterior femur, ventral aspect: 19 Mizotrechus minutus sp. n., ADP 124966; type locality. 20 Mizotrechus dalensi sp. n., ADP 124896; type locality. 21 Mizotrechus chontalesensis sp. n., ADP 127181; type locality. 22 Mizotrechus edithpiafae sp. n., ADP 124948; type locality. 23 Mizotrechus brulei sp. n., ADP 129205; type locality. 24 Mizotrechus gorgona. sp. n., ADP 128622; type locality.

Plate 9. Digital Photo-illustration, male genitalia, median lobe, left lateral, dorsal, and right lateral aspects: 31 Mizotrechus fortunensis sp. n., ADP 124590; type locality. 32 Mizotrechus grossus sp. n., ADP 127165; type locality. 33 Mizotrechus marielaforetae sp. n., ADP 127183; type locality.

Figure 39. Distribution symbol map for known localities of Mizotrechus chontalesensis, Mizotrechus costaricensis, Mizotrechus fortunensis, Mizotrechus gorgona, Mizotrechus jefe, Mizotrechus woldai, spp. n.