Manota vexillifera, Jaschhof & Hippa, 2005

Jaschhof, Mathias & Hippa, Heikki, 2005, The genus Manota in Costa Rica (Diptera: Mycetophilidae), Zootaxa 1011 (1), pp. 1-54 : 31

Manota vexillifera sp. n. ( Fig. 15 )

Description. Small species.

Male. Head. Antenna with fourth flagellomere 1.8 times as long as wide. Maxillary palpus with third palpomere bearing 3, occasionally 4, curved sensilla; fourth palpomere with a parasegment apically.

Thorax. Anterior basalare and laterotergite each bare. Episternum 3 with ventral portion setose. Wing. Length 1.4–1.7 mm. Membrane with a few setae along posterior margin. Sc bare. Junction of CuA2 into CuA1 incomplete.

Terminalia ( Fig. 15 ). Similar to M. montivaga , with differences as follows. Sternite 9 with posterior margin convex medially, situated at mid length of gonocoxites. Disto­lateral lobes of gonocoxites bilobed; on dorsal lobe with large, straight setae pointing posteriorly; on ventral lobe with shorter, straight setae pointing inwards. Gonocoxites in position I with 1 large flag­like megaseta at tip; in position II with 3–4 short, blunt megasetae on lobe; in position III with 2 flat megasetae with one megaseta pointed and the other blunt; in position IV with 1 pointed megaseta, or rarely 2 large setae. Gonostyli small; ventro­apically with 1 large and 1 smaller setae, along dorsal margin with 1+5 curved setae directed dorsally plus 1 isolated seta directed posteriorly.

Diagnostic characters. Among the species with both anterior basalare and laterotergites bare, it has the terminalia with gonocoxites in position I bearing 1 megaseta of unique, flag­like shape.

Etymology. From Latin, vexilliferus, standard­bearing, referring to the peculiar megaseta in position I of the gonocoxites.

Types. Holotype. Male, Costa Rica, Alajuela province, San Carlos, Volcán Arenal National Park , Sendero Pilón , 650 m, 6–7 Nov. 2000, by Malaise trap, G. Carballo ( INBio sample #60084) . Paratypes. 2 males, Cartago province, Tapantí– Macizo de la Muerte National Park, Estación La Esperanza , 2200–2600 m, in cloud forest of oak, 3–22 Nov. 2003, by Malaise trap, M. & C. Jaschhof ; 4 males, Limón province, Pococí, Braulio Carrillo National Park, Estación Quebrada González , 400–500 m, 15 May, 21 Aug. and 24 Sep. 2002, by Malaise trap, P. Hanson & C. Godoy ( INBio samples #68654, 70777 and 71289) .

Other material studied. 2 males, Limón province, Hitoy Cerere Biological Reserve, Sendero Espavel , 11 March–1 April 2003, E. Rojas, B. Gamboa & W. Arana ( INBio sample #73628) ; 1 male, same locality, but 18 Sep.–7 Oct. 2003, E. Rojas, B. Gamboa, W. Arana, M. & C. Jaschhof .

Figures

FIGURE 15. Manota vexillifera sp. n. A: Male terminalia, ventral view (holotype). B: Male terminalia, dorsal view (paratype from Braulio Carrillo). Scale = 0.1 mm.