Manerebia inderena (Adams, 1986)

Mahecha-J, Oscar, Florczyk, Klaudia, Willmott, Keith, Cerdeña, José, Zubek, Anna, Boyer, Pierre, Farfán, Jackie, Lachowska-Cierlik, Dorota & Pyrcz, Tomasz W., 2021, Solving the cryptic diversity of the genus Manerebia Staudinger in northern Peru description of new species and considerations on the biogeographical role of the Huancabamba Deflection (Nymphalidae: Satyrinae: Pronophilina), Zootaxa 5072 (3), pp. 201-237 : 230

Manerebia inderena (Adams, 1986)

Penrosada inderena Adams, 1986: 305 . Type locality: Colombia, Tolima Department, S above Cajamarca. Holotype: NHML [examined].

Remarks. Manerebia inderena has so far been considered as a widely distributed polytypic species. Our preliminary molecular results suggest that at least some of its subspecies should be raised to specific status. The most outstanding case is M. inderena antioquiana Pyrcz & Willmott, 2006 , from the northern portion of the Colombian Central Cordillera, which is highly divergent genetically from other subspecies of M. inderena . Also, M. inderena clara Pyrcz & Willmott, 2006 , from the eastern Andean slopes in central Ecuador, does not segregate with other subspecies of M. inderena . Additionally, M. inderena fina Pyrcz & Willmott, 2006 , from the western slopes in Ecuador, unexpectedly forms a highly supported clade with M. interrupta Brown, 1944 , and M. golondrina Pyrcz & Willmott, 2006 . Finally, M. inderena mirena Pyrcz & Willmott, 2006 , in south-eastern Ecuador is most probably a complex of at least two cryptic species, and further work using a more comprehensive set of samples is underway to clarify the taxonomy of this complex.