Macromesidae new status, 1868

Burks, Roger, Mitroiu, Mircea-Dan, Fusu, Lucian, Heraty, John M., Jansta, Petr, Heydon, Steve, Papilloud, Natalie Dale-Skey, Peters, Ralph S., Tselikh, Ekaterina V., Woolley, James B., van Noort, Simon, Baur, Hannes, Cruaud, Astrid, Darling, Christopher, Haas, Michael, Hanson, Paul, Krogmann, Lars & Rasplus, Jean-Yves, 2022, From hell's heart I stab at thee! A determined approach towards a monophyletic Pteromalidae and reclassification of Chalcidoidea (Hymenoptera), Journal of Hymenoptera Research 94, pp. 13-88 : 13

Macromesidae new status

Macromesinae Graham, 1959. Type genus: Macromesus Walker, 1848.

Diagnosis.

Antenna with 10 flagellomeres in females, 11 in males. Face between malar sulcus and torulus with a second longitudinal sulcus (Fig. 33 ). Clypeus without transverse subapical groove. Labrum subrectangular and hidden, with marginal setae in a row. Mandibles with 3 teeth (Fig. 33 ). Subforaminal bridge with postgena separated by lower tentorial bridge except for a small postgenal bridge dorsal to the hypostoma. Mesoscutellum with frenal arm indicated laterally, and with axillular carina or sulcus. Mesopleural area without an expanded acropleuron. Propodeum with spiracle separated by more than its own length from the anterior propodeal margin (Fig. 34 ). Fore and hind legs with 5 tarsomeres, middle legs in females with 4 tarsomeres. Protibial spur stout and curved; basitarsal comb longitudinal. Metasoma with syntergum, therefore without epipygium.

Discussion.

Macromesus , the only genus of Macromesidae , differs from other Chalcidoidea inthe tarsomere count of females and the usually conspicuous second longitudinal sulcus on the lower face, although it otherwise bears some resemblance to other large-bodied chalcidoids with metallic coloration. The distance from the propodeal spiracle to the anterior margin of the propodeum may cause it to be confused with Ceidae or Hetreulophidae , but this feature is likely convergent, apparently occurring in Macromesidae because of its unusual propodeum.

Figures

Figures 31 - 36. 31, 32 Solenura sp. (Lyscicidae, Solenurinae) 31 antenna 32 mesosoma lateral 33 Macromesus sp. (Macromesidae): head frontal view 34 Macromesus amphiretus Walker (Macromesidae), propodeum 35, 36 Trichilogaster acaciaelongifoliae (Froggatt) (Melanosomellidae) 35 head frontal 36 mesosoma lateral view.