Luponovula maniobraensis, (SQUIRES AND ADVOCATE, 1986)

Groves, Lindsey & Squires, Richard L., 2023, Revision of northeast Pacific Paleogene cypraeoidean gastropods, including recognition of three new species: implications for paleobiogeographic distribution and faunal turnover, PaleoBios 40 (10), pp. 1-52 : 33

LUPONOVULA MANIOBRAENSIS

(SQUIRES AND ADVOCATE, 1986) N. COMB.

FIGS. 9M–P

Eocypraea? maniobraensis Squires and Advocate, 1986 . pp. 856, 857, figs. 2.5, 2.6. Squires, 1991. p. 219, 220, 225; pl. 1, fig. 23.

Eocypraea (Eocypraea) maniobraensis Squires and Advocate. Groves, 1992 . p. 106 [not 1987 as in table 1]. Groves, 1993. p. 11 [not 1983 as in text]. Groves, 1997. p. 8 [as E. (E.) maniobraensis ]. Groves, 2011. p. 46 (table 1).

Eocypraea maniobraensis Squires and Advocate. Fehse, 2001 . p. 20. Groves and Squires, 2021. p. 231 (as Eocypraea ?). Fehse, 2021. p. 42.

Holotype and Type Locality— LACMIP10566.4 , LAC- MIP Type 10566 (ex UCLA 48431 ) ( Figs. 9M–O ), length 39.6 mm, width 22.8 mm, height 21.1. LACMIP Locality 23779 [ex UCLA Locality 3779; ex CSUN Locality 662], Maniobra Formation , Orocopia Mountains , Riverside County, southern California.

Referred Specimen—Hypotype (herein) LACMIP 40491.1 , LACMIP Type 14945 ( Fig. 9P ) , LACMIP Locality 40491 [ex CSUN Locality 491], Llajas Formation, is from a faulted area low in the stratigraphic section of this formation (see geologic map in Squires 1984: fig. 16) in Las Llajas Canyon , north side of Simi Valley , Ventura County, southern California [new information]. This Llajas Formation specimen has been crushed but retains enough morphological features to identify it as L. maniobraensis .

Occurrence— Lower Eocene (“Capay Stage”), Llajas Formation, southwestern Santa Susana Mountains, Los Angeles County, southern California, Ventura County (new information) to Maniobra Formation, Orocopia Mountains, Riverside County, southern California ( Squires and Advocate 1986; Squires 1991).

Etymology— Originally named for the Eocene Maniobra Formation, Riverside County, southern California.

Description— Shell size medium; very inflated; ovatepyriform. Shell widest medially. Base inflated. Spire involute. Aperture narrow and near right margin of shell; aperture elongate anteriorly and curved posteriorly. Aperture narrow and prominently curved. Outer lip narrow, with numerous teeth that become stronger toward anterior of aperture. Inner lip with numerous small teeth.

Remarks— This is an uncommon species. The anteriormost part of the holotype is missing. The dorsal view of this species is shown for the first time in Figure 4M. Luponovula maniobraensis most closely resembles L. oligovata ( Sacco 1894: pl. 3, fig. 25; Cossmann 1903: pl. 7, fig. 2) of late early Oligocene age (Latorfian) from the Liguria and Piedmont regions of Italy and L. normalis ( de Gregorio 1880: pl. 6, fig. 14) of middle Eocene age (Lutetian) from San Giovanni Illarione, Veneto, Italy. In comparison, Luponovula maniobraensis differs by having fewer teeth on its columellar lip, as well as having fewer and shorter teeth on the interior of its outer lip.

Figures

Figure 4 (previous page). A–C. Nucleolaria cowlitzensis Groves, 1994, Cowlitz Formation (middle Eocene, upper “Tejon Stage”), Vader area, south-central Lewis County, Washington. Basal (A), dorsal (B), and left (columellar) side (C) views, 1.9x, length 27.2 mm, width 17.3 mm, height 11.3 mm, of holotype UCMP 39837, UCMP Locality D-8040. D–H. Propustularia kemperae (Nelson, 1925), Santa Susana Formation, (Paleocene, Selandian Stage), north side Simi Hills, Ventura County, southern California. Basal (D), dorsal (E), and right (labral) side (F), and views, 2.2x, length 21 mm, width 17.3, height 12 mm, of holotype CASG 391.01 [ex CASG 987], Locality CASG 391. Dorsal (G) view, 2.6x, length 18.2 mm, width 14.9, height 11.2 mm, of hypotype LACMIP 22307.6, LACMIP Type 14918 (LACMIP Locality 22307). Basal (H) view, 2.2x, length 21 mm, width 17.3, height 12 mm, of paratype UCMP 30541, UCMP Locality 3764. I–N. Subepona goedertorum (Groves and Squires, 1995), Crescent Forma- tion (Eocene, “Capay Stage”), Black Hills, Thurston County, Washington. Basal (I), dorsal (J), left (columellar) lateral (K), and right (labral) (L) views, 2.8x, length 19.4 mm, width 12.3, height 10.8 mm, of holotype LACMIP 16655.23, LACMIP Type 12375, LACMIP Locality 16655 [=LACMIP Locality 41563; ex CSUN Locality 1563]. M–N, Juncal Formation, Canton Canyon, Whitaker Peak area, Ventura County, southern California. Basal (M) and dorsal (N) views, both 2.4x, length 19.4 mm, width 13.9, height 9.4 mm, of hypotype LACMIP 40807.1, LACMIP Type 14935 (LACMIP Locality 40807 [= LACMIP Locality 16191, ex CSUN Locality 807]). O–Q, Subepona leahae n. sp., Llajas Formation (Eocene, “Domengine Stage”), north side Simi Valley, Ventura County, southern California. Basal (O), dorsal (P), and left (labral) side (Q) views, 1.6x, length 29.5 mm, width 13.9, height 15.1 mm, of holotype LACMIP 40374.79, LACMIP Type 14936, (LACMIP Locality 40374).

Figure 9A–B. Grovesia castacensis (Stewart, 1926 [1927]), Tejon Formation (Eocene, “Tejon Stage”), Tehachapi Mountains, Kern County, southern California. Dorsal (A) and left (columellar) (B) views, 3.4x, length 15.5 mm, width 9.1 mm, height 6.9 mm, of hypotype LACMIP 41206.2, LACMIP Type 14941, base of Metralla Sandstone Member, Live Oak Canyon, LACMIP Locality 41206 [ex CSUN Locality 1206]. C–L. Grovesia mathewsonii (Gabb, 1869), Muir Sandstone (Eocene, “Domengine Stage” to “Tejon Stage”), near Martinez, Contra Costa County, northern California. Basal (C), dorsal (D), right (labral) views, 1x, length 10.7 mm, width 7.5 mm, height 4.7 mm, of holotype ANSP 4217. F–G. Basal (F) [as published] and dorsal (G) views, 2.9x, length 17.9 mm, width 11.6 mm, of holotype CASG 245.02 [ex CAS 816] of Cypraea kerniana Anderson and Hanna, 1925, Tejon Formation (Eocene, “Tejon Stage”) Grapevine Canyon, Kern County, southern California. H–J. Basal (H), dorsal (I), left (columellar), (J) left (columellar) views, 2.7x, length 14.6 mm, width 10.8 mm, height 7.8 mm, of hypotype LACMIP 7155.9, LACMIP type 14930, Tejon Formation, Grapevine Canyon, LACMIP Locality 7155, Kern County, southern California. K-L. Basal (K), dorsal (L) views, 3.9x, length 14 mm, width 10.3 mm, of hypotype UCMP 30903, UCMP Locality 5085, Ardath Shale (Eocene, “Domengine Stage”), San Diego County, southern California. M–P. Luponovula maniobraensis (Squires and Advocate, 1986), (M–O). Maniobra Formation (Eocene, “Capay Stage”), Orocopia Mountains, Riverside County, southern California. Basal (M), dorsal (N), and right (labral) (O) views, 1.3x, length 39.6 mm, width 22.8 mm, height 21.2 mm, of holotype LACMIP 10566.4, LACMIP Type 10566 [ex UCLA 48431], LACMIP Locality 23779. (P). Basal view, Llajas Formation (Eocene, “Capay Stage”), north side Simi Valley, Ventura County, southern California, 2.1x, length 26.6 mm, width 15.8 mm, height 10.9 mm, of hypotype LACMIP 40491.1, LACMIP Type 14945 (LACMIP Locality 40491 [ex CSUN Locality 491]).