Lilloiconcha costulata, Hausdorf, 2005

Hausdorf, Bernhard, 2005, The genus Lilloiconcha in Colombia (Gastropoda: Charopidae), Journal of Natural History 39 (30), pp. 2795-2808 : 2796-2802

publication ID	https://doi.org/ 10.1080/00222930500145057
persistent identifier	https://treatment.plazi.org/id/03C78783-FF81-FFE1-FE12-FB07FE3FE26F
treatment provided by	Felipe
scientific name	Lilloiconcha costulata
status	sp. nov.

Treatment

Lilloiconcha costulata sp. n.

( Figures 1 View Figures 1–3 , 4, 5 View Figures 4–9 , 10–15 View Figures 10–12 View Figures 13–20 , 21 View Figure 21 )

Diagnosis

Lilloiconcha costulata differs from other Lilloiconcha species in the depressed conical-globular shell with moderately coarse ribs. The shell is more depressed than in the almost conical Lilloiconcha pleurophora (Moricand, 1846) , but less depressed than in Lilloiconcha gordurasensis ( Thiele, 1927) and the ribs are weaker than in both these species.

Shell ( Figures 1 View Figures 1–3 , 4, 5 View Figures 4–9 )

Depressed conical-globular; with 3.75–4 convex whorls; protoconch with distinct microscopical spiral threads; teleoconch with moderately coarse and more or less regular ribs (ca 6–9 per 1 mm of the body whorl) and a reticulate pattern consisting of fine growthstriae and dense microscopical spiral threads between the ribs; brownish corneous with reddish brown transverse stripes; body whorl rounded; aperture almost circular; upper insertion of the peristome not descending towards the aperture; suture impressed; peristome sharp, neither expanded nor thickened; umbilicus wide, about 24–30% of the shell diameter.

Measurements

La Calera, 0.5 km towards Bogotá (n 515): D52.923.4 mm, Ø53.1¡ 0.2 mm; H51.922.4 mm, Ø52.2¡ 0.2 mm; D/H51.2921.52, Ø51.43¡0.06.

Anatomy ( Figures 10–15 View Figures 10–12 View Figures 13–20 )

One paratype from La Calera, S of Entrada Fabrica de Cemento Samper ( ZMH 4083 View Materials ), has been examined .

Animal greyish; foot aulacopod; tail without distinct caudal-pit, not distinctly truncated; sole not distinctly divided.

Mantle. Mantle with large, fused black spots and small white dots. Mantle collar ( Figure 10 View Figures 10–12 ) broad, with a narrow, indistinctly delimited left mantle lobe and a broad, almost rectangular right mantle lobe. Pneumostome separated by a small, elongate-conical subpneumostomal mantle lobe.

Pallial complex ( Figure 11 View Figures 10–12 ). Pallial complex about three times as long as the broad kidney, which extends to the hindgut. The kidney is slightly longer than the pericardium. It opens into the reflexed primary ureter near its anterior end. The primary ureter is continued by a tube-like secondary ureter along the hindgut, which ends with the ureteric pore opening next to the anus inside the pneumostome. Lung long and narrow, without coarse venation.

Genitalia ( Figure 12 View Figures 10–12 ). The male copulatory organs are strongly reduced. The penis forms a lateral swelling at the large atrium and contains a conical papilla. The penis retractor inserts near the proximal end of the penis and at the diaphragm. It crosses the right ommatophoral retractor. The vas deferens is very broad and enters the penis terminally. No differentiated epiphallus could be discerned. The vagina is very short. The peduncle of the bursa copulatrix is long. The small bursa of the bursa copulatrix reaches the digestive gland. The free oviduct is short. The right ommatophoral retractor does not run between penis and vagina.

Radula ( Figures 13–15 View Figures 13–20 ). The tricuspid central teeth are narrower and shorter than the adjacent lateral teeth. The slender central cusps of the central teeth are about three times as long as the side cusps. The central cusp of the tricuspid lateral teeth is shifted backwards. It is connected with the large and strongly curved side cusps by a low bridge which begins between the bases of the side cusps with an apparent extra denticle. The side cusps of the lateral teeth are raised above the elevation plane of the mesocone. Towards the periphery the endocone of the lateral teeth increases and approximates the mesocone. At the transition to the marginal teeth extra denticles develop next to the ectocone so that the marginal teeth are multicuspid.

Etymology

The specific epithet refers to moderately coarse ribs which characterize the species (Latin costulatus).

Type locality

Colombia, Cundinamarca: La Calera, 0.5 km towards Bogota´, farm Las Cabreras, Vereda San Rafael, Andean forest , N slope, 2840 m altitude, 04 ° 429300N, 73 ° 589200W .

Distribution ( Figure 21 View Figure 21 )

Lilloiconcha costulata is known from Andean forests at 2400–3100 m altitude in the Distrito Especial and the Departamentos Cundinamarca and Boyacá in Colombia .

Material

Colombia, Distrito Especial: Bogota´, forest near brook above Escuela Militar de Caballeria , 2750 m altitude, 04 ° 419150N, 74 ° 019330W ( ZMH 4322 View Materials ) . Colombia, Cundinamarca: La Calera, 0.5 km towards Bogota´, farm Las Cabreras, Vereda San Rafael , 2820–2840 m altitude, 04 ° 429300N, 73 ° 589200W (holotype ZMH 4793 View Materials , leg. B. Hausdorf, 11 March 2000, measurements: D 53.2 mm, H 52.2 mm; UNAL; ZMH 4021 View Materials , 4037 View Materials , 4069 View Materials ) ; La Calera, S of Entrada Fabrica de Cemento Samper, degraded Andean forest , NW slope, 2750 m altitude, 04 ° 439150N, 73 ° 569460W ( ZMH 4083 View Materials ) ; Bosques de Torca, Andean forest , W slope, 2800 m altitude, 04 ° 499180N, 74 ° 019070W ( ZMH 4334 View Materials ) ; Laguna Guatavita, Andean forest , S slope, 3030 m altitude, 04 ° 589450N, 73 ° 469240W ( ZMH 4096 View Materials ) ; Bogotá towards Tunja, Alto del Sisga, Andean forest , N slope, 2890 m altitude, 05 ° 049580N, 73 ° 449420W ( ZMH 4103 View Materials ) . Colombia, Boyaca´: Arcabuco 21 km towards Tunja, Alto del Sote , Andean forest , 3100 m altitude, 05 ° 399N, 73 ° 229W ( ZMH 4240 View Materials ) ; Moniquira 13.2 km towards Arcabuco , waterfall above road, 2400 m altitude, 05 ° 499230N, 73 ° 309310W ( ZMH 4142 View Materials ) .