Lantana undulata Schrank (1822:56)

Cardoso, Pedro Henrique, Valério, Vanessa Imaculada Dos Reis, Neto, Luiz Menini & Salimena, Fátima Regina Gonçalves, 2021, Verbenaceae in Espírito Santo, Brazil: richness, patterns of geographic distribution and conservation, Phytotaxa 484 (1), pp. 1-43 : 19-20

6.10. Lantana undulata Schrank (1822:56) . (Figs. 2B, 6F)

Shrubs 1‒1.2 m high, branches tetragonal, without prickles, sparsely strigose, glandular sessile trichomes present. Leaves opposite, petiole 3‒5 mm long; blade 4.3‒9.5 × 2‒4.3 cm, chartaceous, elliptical, apex acute, base cuneate, margin entire near the base, crenate toward the apex, ciliate, adaxial surface opaque, strigose, slightly bullate, abaxial surface strigose along with the veins or tomentose along the entire blade, glandular sessile trichomes present. Inflorescences 1‒1.5 cm long, rachis elongated up to 6.5 cm long in fructification, 1‒2 per axil, peduncle 3.7‒8.5 cm long, sparsely strigose, glandular sessile trichomes present; bracts unequal, external 0.9‒1 cm long, internal 7‒8 mm long, wide-ovate to ovate, abaxial surface sparsely strigose, glandular sessile trichomes present, margin ciliate; calyx 1.7‒2 mm long, truncate, sparsely strigose externally, glandular sessile trichomes present, accrescent in fruit; corolla 0.8‒1 cm long, white, throat yellow. Fruit ca. 3 mm long, external surface rugose, vinaceous to black, surrounded by the persistent calyx.

Distribution and habitat:— Lantana undulata is endemic to Brazil, distributed from Paraíba to Santa Catarina ( Silva & Lima 2012, Silva et al. 2020). In Espírito Santo, it occurs in restinga and dense rainforest, and often found in disturbed areas. Not collected in SPAs in this state.

Phenology:— Collected with flowers and fruits in January, April, May, June, July, and August.

Preliminary conservation assessment: —This is a common species with a wide distribution ( Silva & Lima 2012), and occurs inside PAs in Brazil. Considered “Least Concern” (LC) ( IUCN 2019).

Selected material:— BRAZIL. Espírito Santo: Aracruz, 6 August 2018, fl. and fr., D. T . Wandekoken & G. G . Ruschi 333 ( VIES); Conceição da Barra, Floresta Nacional do Rio Preto , 9 April 2011, fl. and fr., T. L . Rocha & M . Ribeiro 117 ( CESJ, SAMES); Ibiraçu , 27 May 1990, fl. and fr., H. Q. B . Fernandes 2939 ( CESJ, MBML, SPF, VIES); Linhares , 10 May 1977, fl. and fr., G . Martinelli 1867 ( SPF); Nova Venécia , 12 January 1953, fl. and fr., A. P . Duarte 3726 ( RB); Serra , 4 April 1992, fl. and fr., R. G . Silveira s.n. ( CESJ 26076 ); Santa Teresa, 29 May 1984, fl. and fr., W . Boone 204 ( CESJ, MBML); São Mateus , 7 August 1983, fl. and fr., G . Hatschbach 46794 ( CEPEC, MBM); Sooretama , 13 May 2009, fl., G . D. Colletta et al. 305 ( ESA, RB, VIES); Vitória , 3 July 1984, fl. and fr., N. M. S . Varejão s.n. ( CESJ 65202 , VIES 501 ) .

Notes:— Lantana undulata is characterized by the branches without prickles; elliptical leaves; rachis elongated up to 6.5 cm long in fructification; bracts sparsely strigose in the abaxial surface; truncated calyx, and white corolla.

Illustrations in Silva & Lima (2012).