Laccosmylus latizonus, Fang, Hui, Ren, Dong, Liu, Jiaxi & Wang, Yongjie, 2018

Fang, Hui, Ren, Dong, Liu, Jiaxi & Wang, Yongjie, 2018, Revision of the lacewing genus Laccosmylus with two new species from the Middle Jurassic of China (Insecta, Neuroptera, Saucrosmylidae), ZooKeys 790, pp. 115-126 : 120-123

Laccosmylus latizonus sp. n. Figures 4, 5

Diagnosis.

Forewing with four pigmented patch-like markings. Hind wing with three pigmented stripes, and the third stripe interlinked with a fuscous patch covering wing apex. The venation of fore- and hind wing: RA area forming 4-5 rows of irregular cells; 1 to 2 rows of cells between longitudinal veins from radius area to anal area.

Description.

Forewing more than 54.4 mm long, partially preserved, outer margin unknown (Figure 4A, B). Four pigmented patch-like markings present. Costal veinlets forked twice distally, interlinked by 5-7 rows of smaller veinlets. ScP fused with RA apically. RA-RP area broad, forming 4-5 rows of irregular cells. RP sharply bent towards RA terminally, forming an angle of approx. 45°. Single row of cells between main longitudinal veins from radius area to anal area. MA and MP partially preserved. CuA forming a large triangular area.

Hind wing length 56-59 mm, width 22-26 mm. Hind wing broad, with outer margin slightly undulant, same shape as the type species L. calophlebius (Figures 1A, 5A), covered with 3 pigmented stripes, and the third stripe interlinked with a fuscous patch covering wing apex. (Figure 5A, C). Trichosors present along distal half of wing margin. Hind wing showing similar vein pattern as forewing from costal area to media area (Figures 4, 5A, B). In addition, RP1 deeply branched at the base or not (Figure 4A, B, D); MA forming three main branches distally; MP forked before the separation of MA from RP; MP forming several pectinate branches distally; CuP with several pectinate branches. AA1 and AA2 simple, partially preserved.

Type material.

Holotype: CNU-NEU-NN2018008P/C, only hind wing preserved; Paratypes: CNU-NEU-NN2018009, four wings partially preserved and joined together; CNU-NEU-NN2018010P/C, only hind wing partially preserved. These specimens are deposited in the Key Laboratory of Insect Evolution and Environmental Changes, College of Life Sciences, Capital Normal University, Beijing, China.

Type locality and horizon.

Jiulongshan Formation, Daohugou locality (41°18.5'N, 119°13'E (DDM)), Shantou Township, Ningcheng County, Chifeng City, Inner Mongolia, China; Middle Jurassic, Bathonian-Callovian boundary.

Etymology.

The Latin latizonus is derived from the stripe-like wing markings of this species.

Remarks.

The new species evidently belongs to Laccosmylus according to the emended diagnosis of the genus, i.e., the same broad hind wing shape, undulate outer margin and similar venation, and even the arrangement of wing markings. Laccosmylus latizonus sp. n. can be easily distinguished from L. calophlebius and L. cicatricatus sp. n. by the following characters of the hind wing markings, i.e. three distinct pigmented stripes in the L. latizonus vs. scattered patch-like markings on the other two species. In addition, the venation of these species are also distinctively different, e.g., L. latizonus sp. n. with 4-5 rows of irregular cells between RA and RP vs. 6-7 rows of irregular cells in the other two species; relatively simpler cross-veins in L. latizonus sp. n. in radial sector than in the other two species.