Incasarcus dianae Kury & Maury, 1998

Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo, 2022, Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species, Arthropod Systematics & amp; Phylogeny 80, pp. 309-388 : 309

3.32. Incasarcus dianae Kury & Maury, 1998

Figs 4C , 9C, D , 13G, H , 30

Incasarcus dianae Kury & Maury, 1998: 146 (desc.), 159 (key), figs. 1 (male dorsal habitus, chelicerae, pedipalpus, trochanter-patella IV), 2 (penis dorsal view), 3 (penis lateral view), 4 (male lateral habitus), 5 (female dorsal habitus, chelicerae, pedipalpus), 6 (male ventral habitus); 7-10 (tarsi I-IV); Kury 2003: 144 (cat.), Pinto-da-Rocha et al. 2014: 6 (cit.); Kury 2014: 7 (cit.), 11 (cit.), 47 (mat.); Kury 2016: 146 (cit.), figs. 3a (penial microsetae).

Redescription.

MALE: Measurements (n =3) DSW: 4.0-4.3 (4.0); DSL: 4.5-5.1 (4.5); CL: 1.1-1.5 (1.1). FIVL: 11.3-11.4 (11.4). ChL: 3.3-3.4 (3.4). Coloration (in ethanol): Yellow with black spots covering practically the entire dorsal scutum; area III brown. Pedipalpus , chelicerae and legs I-IV brown. Dorsum: (Fig. 4C ) Anterior margin of dorsal scutum granulate. Ocularium with well-defined median depression; sparsely granulate. Carapace sparsely granulate. Areas I-IV of dorsal scutum with scattered granules, with a slightly higher density at areas I-II. Area I with a pair of small lateral tubercles, slightly larger than granules. Area II and IV unarmed. Area III with a median pair of spines, directed backwards. Lateral margins of dorsal scutum with scattered granules. Posterior margin of dorsal scutum and free tergites I-III almost smooth, with few granules sparsely distributed. Free tergites I-III with a pair of median acuminate tubercles. Chelicerae: (Fig. 4C ) Swollen. Segment I with few granules distally. Segment II covered with small granules; with eight teeth. Segment III with five teeth. Pedipalpus : Trochanter with a ventroapical setiferous tubercle. Femur with a ventrobasal setiferous tubercles; a ventral row of three setiferous tubercles; with a large proapical spine. Patella with a proapical tubercle. Venter: Coxa I with a median row of 4-5 setiferous tubercles; II-IV granulate. Rows of small tubercles between the coxae II-III and III-IV. Stigmatic area slightly granulate. Free sternites I-III with a row of small granules. Anal operculum granulate. Legs: (Figs 4C , 9C, D ) Coxae I-II each with a prolateral and a retrolateral apophysis. Coxa III with a prolateral apophysis. Coxa IV with scattered setiferous granules. Trochanters I-III granulate and unarmed. Trochanter IV granulate and with a retrolateral tubercle. Femora I-III with sparsely distributed small granules. Femur IV covered with small granules; a retrolateral row of 22-24 acuminate tubercles along basal ⅔, decreasing in size apically; a prolateral a row of 28-30 acuminate tubercles along basal ⅔, decreasing in size apically. Patellae I-III unarmed. Patella IV with a pair of dorsoapical tubercles (retrolateral one larger.) Tibiae I-IV with few scattered granules. Tarsal segmentation: (n =3) 9, 15-17 (17), 10, 11. Penis: (Fig. 13G, H ) VP rectangular; with straight distal margin, with conspicuous lateroapical projections. MS C1-C4(C5) subapical long and apically curved; MS A1-A2 median short and straight; MS D1 very short, dorsally placed near MS C; MS E1-E2(E3) very short, placed in lateral flanges of VP. Lateral sacs short, with blunt apex; with T3-like short microsetae. Stylus with broad apex; apically with small projections. Promontory truncated. - FEMALE: Measurements (n =3) DSW: 4.4-5.0; DSL: 5.5-5.8; CL: 1.3-1.4; FIVL: 11.2-11.5. ChL: 1.3-1.4. Females more robust than the males. Chelicerae not swollen. Femur IV unarmed. Tarsal segmentation: (n =3) 8-9, 14-16, 10-11, 11-13.

Diagnosis.

It differs from other species of the genus by the set of following characters: DS without silver-white coloration; DS slightly granulate; area I with a pair of lateral tubercles; area III with a pair of spines (Fig. 4C ); male femur IV with a retrolateral and a prolateral rows of tubercles at ⅔ basal (Fig. 9C, D ); penis with short lateral sacs (Fig. 13G, H ).

Distribution.

(Fig. 30 ) PERU. Cusco. Paucartambo province, Manu National Park.

Material examined.

Type material: Holotype ♂, ' PERU, Cusco, Paucartambo province, Manu National Park, road to Paucartambo-Pilcopata , 2,900 m a.s.l., 13°01′40″S 71°16′40″W | 19/II/1990, A. Cano & D. Silva leg. (MUSM 410) - Paratypes 2 ♀, ‘ditto’ (MUSM 410); Paratypes 1 ♂, 2 ♀, ' PERU, Cusco, Paucartambo province, Manu National Park,| road to Paucartambo-Pilcopata , 2,900 m, 13°01′40″S 71°16′40″W, 14/II/1990, A. Cano & D. Silva leg. (MNRJ 5315 ex- MUSM 410) .

Figures

Figure 4. Habitus, dorsal of males of Huancabamba gen. nov. and Incasarcus. A H. kubricki gen. et sp. nov.; B I. argenteus Kury & Maury, 1998; C I. dianae Kury & Maury, 1998; D I. ochoai Kury & Maury, 1998; E I. pictus Kury & Maury, 1998; F I. viracocha Kury & Maury, 1998; Legend bars = 1 mm.

Figure 9. Male trochanter-patella / tibia IV of Incasarcus. A - B Trochanter-tibia IV of I. argenteus Kury & Maury, 1998, dorsal and ventral view, respectively; C - D Trochanter-tibia IV of I. dianae Kury & Maury, 1998, dorsal and ventral view, respectively; E-F Trochanter-tibia IV of I. pictus Kury & Maury, 1998, dorsal and ventral view, respectively; G - H Trochanter-tibia IV of I. viracocha Kury & Maury, 1998, dorsal and ventral view, respectively; I Trochanter-patella of I. viracocha, lateral view; J Trochanter-patella of I. ochoai Kury & Maury, 1998, large male, dorsal view; K - L Trochanter-tibia of I. ochoai, small male, dorsal and ventral view, respectively; M Trochanter-patella of I. ochoai, large male, lateral view; N Trochanter-patella of I. ochoai, small male, lateral view; Legend bars = 1 mm.

Figure 13. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B A. vargasllosai sp. nov.; C - D A. weyrauchi (Roewer, 1952) comb. nov.; E - F Incasarcus argenteus Kury & Maury, 1998; G - H Incasarcus dianae Kury & Maury, 1998; I - J: Incasarcus pictus Kury & Maury, 1998; Legend bars = 0.1 mm.

Figure 30. Distribution of Incasarcus in Cusco, Peru.