Hoploscopa subvariegata (Rotschild, 1915)

Leger, Theo, Kehlmaier, Christian, Vairappan, Charles S. & Nuss, Matthias, 2020, Twenty-six new species of Hoploscopa (Lepidoptera, Crambidae) from South-East Asia revealed by morphology and DNA barcoding, ZooKeys 907, pp. 1-99 : 1

Hoploscopa subvariegata (Rotschild, 1915) Figs 25 , 64 , 109

Material examined.

Holotype: ♀, with labels: "Holo- | type" [round label, red ringed]; "Angabunga R[iver]., | affl[uent]. of St. Joseph | R., Brit[ish].N[ew].Guinea, | 6000 f[ee]t, upwards. | Nov.[19]04.-Febr.[19]05. | (A. S. Meek)."; "Eudorina | subvariegata | Type Rotsch[ild]." [handwritten]; "♀ | Pyralidae | Brit[ish].Mus[eum]. | Slide N°. | 20254"; " NHMUK 010923326" [barcode appended]. Deposited in NHMUK.

Other specimens examined.

14 ♂, 4 ♀. Papua New Guinea: 10 ♂ (2 with genitalia on slide TL478 ♂ & TL642 ♂), 1 ♀ (genitalia on slide TL521 ♀), Morobe Province, near Wau, Mt. Kaindi, 2360 meters, 27-28 July 1983, leg. S. E. & P. M. Miller; 1 ♂ (genitalia on slide TL534 ♂), same data except 29-30.viii.1983; 1 ♂, 1♀, same locality, 11.xii.1976, leg. G. F. Hevel & R. E. Dietz; 2 ♂ (1 with DNA voucher 3156, genitalia on slide TL436 ♂), 1 ♀, same locality, 3.x.1992, leg. V. O. Becker); 1 ♂, 1 ♀ (genitalia on slide TL471 ♀), Morobe Province, Biaro Road, 2000 m, 25.ix.1992, leg. V. O. Becker (USNM).

Diagnosis.

Hoploscopa subvariegata displays four roughly triangular yellow spots filled with reddish brown on forewing costa. In male genitalia, uncus is long and slender with truncate apex, gnathos projection is tongue-shaped, reaching 1/3 of uncus length, and dorsal margin of the valva is conspicuously protruded dorsad. In female genitalia, ductus bursae is long and straight, corpus bursae is large, globular and bears a small, slightly curved thorn with leaf-shaped sclerotisation at its base.

Similar species.

Hoploscopa persimilis . The latter species lacks the subterminal triangular yellow blotch at forewing costa. In the male genitalia of H. persimilis , uncus apex is duck-shaped, gnathos projection forms a small triangular tip, and valva dorsal margin is not protruded. Female of H. persimilis is not known.

Description.

Head. Antennae dorsally with bronze scales. Proboscis pale yellow, basally brown. Maxillary palpi brown. Labial palpi brown, ventro-basally pale yellow.

Thorax (Fig. 25 ). Collar white. Forewing length: 10 mm (♂), 9.5-12 mm (♀); forewing ground colour brown to dark brown; basal area reddish brown, basally with small yellow dash, at costa with V-shaped yellow patch filled with reddish brown; antemedian cubital snow-white dot; costal field reddish brown; median discoidal stigma V-shaped, white, yellow toward costa, filled with reddish brown; postmedian patch reddish brown, with white to yellow basal edge and pale yellow to yellow costal blotch; subterminal costal spot pale yellow to yellow, encroached with reddish brown; subterminal field reddish brown; margin brown, with white dots. Hindwing dirty white, bronze toward margin. Forelegs femur and tibia brown; tarsi pale yellow. Midlegs with femur brown; tibia pale yellow, dorso-distally brown; tarsi pale yellow. Hindleg femur brown; tibia pale yellow, dorso-distally brown; tarsi pale yellow to pale brown.

Abdomen. Male sternum A8 posterior margin bilobed.

Male genitalia (N = 4) (Fig. 64 ). Uncus long and slender, gently tapering toward apex, apex roughly truncate. Gnathos projection tongue-shaped, ca. 1/3 of uncus length. Valva ventral margin nearly straight, dorsal margin strongly protruded dorsad, apex truncate. Juxta with broad rounded base, medially wide, apex broadly incurved, weakly sclerotised. Saccus broad, triangular, pointing dorsad. Phallus with elongated, flat, spatula-shaped cornutus apically with narrow bump.

Female genitalia (N = 2) (Fig. 109 ). Anterior apophyses with small dorsal tip at posterior 1/3. Antrum sclerotisation short, as long as broad. Ductus bursae of medium length, slender, straight. Corpus bursae large, globular, posterior half reticulated, anterior half membranous, with sclerotisation between thorn and corpus opening, medially with faintly marked sclerotised band. Thorn slightly curved, with small dents pointing toward thorn base.

Distribution.

Recorded from the Angabunga River in the Central Province (Papua New Guinea), also known from Mount Kaindi, Morobe Province, at altitudes between 2,000 m and 2,360 m.

Remarks.

Nuss (1998) transferred this species from Eudorina to Hoploscopa . Other specimens examined from Papua New Guinea display the same pattern and identical genitalia as the holotype from Western New Guinea.

Figures

Figures 19 - 33. Forewing of Hoploscopa species 19 Hoploscopa danaoensis sp. nov., paratype, ♂, Philippines, Leyte, Lake Danao, 650 m, 14 - 17. iv. 1997 (Mey & Speidel) 20 Hoploscopa aurantiacalis Snellen lectotype, ♀, NHMUK 010923286, Indonesia, Java occ., Pengaleng, 4000 ft, 1893 (genitalia on slide Pyralidae Brit. Mus. Slide N ° 20246) 21 Hoploscopa brunnealis Snellen, lectotype, ♀, NHMUK 010923292, Indonesia, Java occ., Pengaleng, 4000 ft, 1893 (genitalia on slide Pyralidae Brit. Mus. Slide N ° 20247) 22 Hoploscopa ocellata Hampson, holotype, ♀, NHMUK 010923358, Indonesia, Moluccas, Batchian, iii. 1892 (W. Doherty) (genitalia on slide Pyralidae Brit. Mus. Slide N ° 20258) 23 Hoploscopa quadripuncta Rotschild, holotype, ♂, NHMUK 010923357, Indonesia, Papua, Utakwa River, 3000 ft, i. 1913 (A. F. R. Wollaston) (genitalia on slide Pyralidae Brit. Mus. Slide N ° 20257) 24 Hoploscopa semifascia Hampson, ♀, NHMUK 010923321, Papua New Guinea, Southern Highlands, Bosavi, 570 m, 6. i. 1986 (D. Agassiz) (genitalia on slide TL 740 ♀) 25 Hoploscopa subvariegata Rothschild, ♂, Morobe Province, near Wau, Mt. Kaindi, 2360 m, 27 - 28. vii. 1983 (S. E. & P. M. Miller) (genitalia on slide TL 642 ♂) 26 Hoploscopa persimilis Rothschild, lectotype, ♂, NHMUK 010923328, Indonesia, Papua, Utawka River, 3000 ft., i. 1913 (A. F. R. Wollaston) (genitalia on slide Pyralidae Brit. Mus. Slide N ° 20255) 27 Hoploscopa astrapias Meyrick, lectotype, ♀, NHMUK 010923383, Fiji, Vunidawa, 2. i. 1932 (genitalia on slide Pyralidae Brit. Mus. Slide N ° 20241) 28 Hoploscopa anamesa Tams, ♀, NHMUK 010923466, Vanuatu, Aneityum Island, Red Crest, 3 m NE of Anelgauhat, 1200 ft, vi. 1955 (E. Cheesman) (genitalia on slide TL 718 ♀) 29 Hoploscopa nauticorum Tams, allotype, ♀, Samoan, Upolu Island, Malololelei, 2000 ft, 21. ii. 1925 (P. A. Buxton & G. H. Hopkins) 30 Hoploscopa diffusa Hampson, lectotype, ♂, NHMUK 010923338, Papua New Guinea, Fergusson Island, x-xi. 1894 (A. S. Meek) (genitalia on slide Pyralidae Brit. Mus. Slide N ° 1014 ♂) 31 Hoploscopa triangulifera Hampson, ♀, NHMUK 010923453, Biagi, Mambare R., 1600 m, iii. 1906 (A. S. Meek) (genitalia on slide TL 712 ♀) 32 Hoploscopa anacantha sp. nov., holotype, ♂, NHMUK 010923444, Indonesia, North Sumatra, Dumoga-Bone N. P., Gunung Mogogonipa, summit, 1008 m, 18 - 20. x. 1985 (genitalia on slide TL 759 ♂) 33 Hoploscopa kelama sp. nov., paratype, ♂, NHMUK 010923342, Indonesia, North Sulawesi, Dumoga-Bone National Park, Clarke Camp, lower montane forest, 1140 m, x. 1985 (Royal Entomological Society of London, Project Wallace) (genitalia on slide TL 759 ♂).

Figures 64 - 69. male genitalia features of Hoploscopa. 64 Hoploscopa subvariegata, TL 534 ♂ 65 Hoploscopa persimilis, lectotype, Pyralidae Brit. Mus. Slide N ° BMNH 20255 ♂ (no juxta drawing) 66 Hoploscopa diffusa, holotype, Pyralidae Brit. Mus. Slide N ° BMNH 1014 ♂ 67 Hoploscopa triangulifera, TL 711 ♂ 68 Hoploscopa astrapias, TL 724 ♂ 69 Hoploscopa anamesa, TL 727 ♂.

Figures 107 - 110. Female genitalia features of Hoploscopa. 107 Hoploscopa astrapias, TL 23 ♀ 108 Hoploscopa anamesa, TL 718 ♀ 109 Hoploscopa subvariegata, lectotype, Pyralidae Brit. Mus. Slide N ° BMNH 20254 ♀ 110 Hoploscopa triangulifera, holotype, Pyralidae Brit. Mus. Slide N ° BMNH 20256 ♀.