Homoneura (Homoneura) degenerata, Shi, Li & Yang, Ding, 2014

Shi, Li & Yang, Ding, 2014, Supplements to species groups of the subgenus Homoneura in China (Diptera: Lauxaniidae: Homoneura), with descriptions of twenty new species, Zootaxa 3890 (1), pp. 1-117 : 54-55

Homoneura (Homoneura) notostigma group

88. Homoneura (Homoneura) notostigma (Kertész, 1913) . Oriental: China (Hainan, Taiwan), Thailand.

Homoneura (Homoneura) ornatifrons group

89. Homoneura (Homoneura) ornatifrons (Kertész, 1913) . Oriental: China ( Taiwan), Japan (Ryukyu), Vietnam. ( Figs. 179–183 ).

Homoneura (Homoneura) picta group

90. Homoneura (Homoneura) picta (Meijere, 1904) . Palaearctic: Moscow; Oriental: China (Guangxi, Guizhou, Hainan, Taiwan, Zhejiang), India, Indonesia, Malaysia, Nepal, Thailand, Vietnam.

91. Homoneura (Homoneura) procerula Gao & Yang, 2005 . Oriental: China (Guizhou).

92. Homoneura (Homoneura) trypetoptera (Hendel, 1908) . Oriental: China (Guizhou, Hainan, Taiwan), India, Indonesia, Laos, Malaysia, Nepal, Philippines, Sri Lanka, Thailand, Vietnam. ( Figs. 315–318 ).

Homoneura (Homoneura) quinquenotata group

93. Homoneura (Homoneura) brevicornis (Kertész, 1915) . Oriental: China (Heilongjiang, Hainan, Taiwan), Indonesia, Phillippines, Japan (Ryukyus), Australian/Oceanian: New Guinea, Solomon Islands.

94. Homoneura (Homoneura) cangshanensis Li & Yang, 2014 . Oriental: China (Yunnan).

95. Homoneura (Homoneura) crispa Li & Yang, 2014 . Oriental: China (Yunnan).

96. Homoneura (Homoneura) haejuana Sasakawa & Kozánek, 1995 . Palaearctic: China ( Tianjin), North Korea.

97. Homoneura (Homoneura) levis (Wiedemann, 1830) . Oriental: China ( Macao). ( Fig. 13 )

98. Homoneura (Homoneura) litorea Shi, Gaimari & Yang, 2012 . Oriental: China (Hainan).

99. Homoneura (Homoneura) longinotata Shi & Yang, 2009 . Oriental: China (Hainan).

100. Homoneura (Homoneura) quinquenotata (Meijere, 1915) . Oriental: China (Fujian, Hainan, Taiwan), Indonesia, Laos, Malaysia, Vietnam. ( Figs. 205–216 ).

101. Homoneura (Homoneura) semiannulata Li & Yang, 2014 . Oriental: China (Yunnan).

102. Homoneura (Homoneura) stackelbergiana Papp, 1984 . Palaearctic: China (Beijing), North Korea, Russia.

103. Homoneura (Homoneura) suturalis Yang, Zhu & Hu, 2003 . Oriental: China (Fujian).

104. Homoneura (Homoneura) trisurstylata Li & Yang, 2014 . Oriental: China (Yunnan).

Homoneura (Homoneura) quinquevittata group

105. Homoneura (Homoneura) interrupta Sasakawa, 2001 . Oriental: China (Hainan), Vietnam. ( Figs. 10 , 138–142 ).

106. Homoneura (Homoneura) quinquevittata (Meijere, 1910) . Oriental: China (Fujian, Guizhou, Hainan, Taiwan, Yunnan), India, Indonesia, Japan (Ryukyu), Laos, Malaysia, Nepal, Philippines, Thailand. ( Figs. 217–223 ).

107. Homoneura (Homoneura) strigata (Meijere, 1910) . Oriental: China (Guangxi, Hainan), Indonesia, Malaysia, Laos, Vietnam. ( Figs. 272–283 ).

Homoneura (Homoneura) sauteri group

108. Homoneura (Homoneura) conspicua Sasakawa, 2001 . Oriental: China (Hainan), Vietnam. ( Figs. 2 , 76–80 ).

109. Homoneura (Homoneura) degenerata sp. nov. Oriental: China (Hainan). ( Figs. 4 , 97–101 ).

110. Homoneura (Homoneura) longiplumaria Yang, Hu & Zhu, 2002 . Oriental: China (Hainan).

111. Homoneura (Homoneura) sauteri Malloch, 1927 . Oriental: China (Hainan, Taiwan), Laos, Malaysia, Sri Lanka, Vietnam. ( Figs. 224–234 ).

112. Homoneura (Homoneura) septentrionalis (Loew, 1847) . Palaearctic: China (Harbin), Mongolia, Russia. ( Figs. 235–243 ).

113. Homoneura (Homoneura) simplicissima (Meijere, 1910) . Oriental: China ( Taiwan), Indonesia, Malaysia, Philippines. ( Figs. 260–271 ).

Homoneura (Homoneura) singularis group

114. Homoneura (Homoneura) aulatheca Sasakawa & Ikeuchi, 1985 . Palaearctic: Japan, North Korea; Oriental: China ( Taiwan).

115. Homoneura (Homoneura) concava Sasakawa, 2002 . Oriental: China ( Taiwan). ( Figs. 71–75 ).

116. Homoneura (Homoneura) didyma Yang, Zhu & Hu, 2003 . Oriental: China (Fujian).

117. Homoneura (Homoneura) falcata sp. nov. Oriental: China (Hainan). ( Figs. 6 , 107–111 ).

118. Homoneura (Homoneura) fengyangshanica sp. nov. Oriental: China (Zhejiang). ( Figs. 7 , 112–116 ).

119. Homoneura (Homoneura) hongmaoensis sp. nov. Oriental: China (Hainan). ( Figs. 9 , 133–137 ).

120. Homoneura (Homoneura) latifrons Malloch, 1927 . Oriental: China ( Taiwan), Japan (Kyushu, Ryukyus). ( Figs. 152–157 ).

121. Homoneura (Homoneura) longicornis Sasakawa, 2002 . Oriental: China ( Taiwan).

122. Homoneura (Homoneura) singularis Yang, Hu & Zhu, 2002 . Oriental: China (Hainan).

123. Homoneura (Homoneura) subvittata Malloch, 1927 . Oriental: China ( Taiwan), Indonesia, Malaysia, Thailand. ( Figs. 289–299 ).

124. Homoneura (Homoneura) tianlinensis Gao & Yang, 2004 . Oriental: China (Guangxi).

125. Homoneura (Homoneura) zhejiangensis sp. nov. Oriental: China (Zhejiang). ( Figs. 24 , 332–336 ).

126. Homoneura (Homoneura) vittigera Sasakawa, 2001 . Oriental: China (Hainan, Yunnan), Vietnam.

Homoneura (Homoneura) striatifrons group

127. Homoneura (Homoneura) chishuiensis Gao & Yang, 2006 . Oriental: China (Guizhou, Hainan, Yunnan).

128. Homoneura (Homoneura) shunxica sp. nov. Oriental: China (Zhejiang). ( Figs. 18 , 244–248 ).

Homoneura (Homoneura) substigmata group

129. Homoneura (Homoneura) substigmata Yang, Zhu & Hu, 1999 . Oriental: China (Henan).

Homoneura (Homoneura) tibetensis group

130. Homoneura (Homoneura) lata Yang, Hu & Zhu, 2001 . Oriental: China (Zhejiang).

131. Homoneura (Homoneura) pictipennis Czerny, 1932 . Palaearctic: China (Harbin), Russia. ( Figs. 190–194 ).

132. Homoneura (Homoneura) polyacantha Yang, Zhu & Hu, 1999 . Oriental: China (Henan).

133. Homoneura (Homoneura) tibetensis Gao & Yang, 2003 . Palaearctic: China (Tibet).

134. Homoneura (Homoneura) variinervis (Kertész, 1915) . Palaearctic: Japan (Honshu). Oriental: China ( Taiwan), Japan (Ryukyu), Thailand.

Homoneura (Homoneura) trispina group

135. Homoneura (Homoneura) bicuspidata sp. nov. Oriental: China (Hainan). ( Figs. 1 , 34–38 ).

136. Homoneura (Homoneura) noticomata sp. nov. Oriental: China (Hainan). ( Figs. 15 , 163–167 ).

137. Homoneura (Homoneura) trispina Malloch, 1927 . Oriental: China (Hainan), Indonesia, Malaysia, Singapore.

Homoneura (Homoneura) unguiculata group

138. Homoneura (Homoneura) diversa (Kertész, 1913) . Oriental: China (Fujian, Taiwan), Philippines.

139. Homoneura (Homoneura) jianlingensis sp. nov. Oriental: China (Hainan). ( Figs. 11 , 143–146 ).

140. Homoneura (Homoneura) pallidula Malloch, 1927 . Oriental: China ( Taiwan). ( Figs. 184–189 ).

141. Homoneura (Homoneura) unguiculata (Kertész, 1913) . Palaearctic: Japan (Honshu, Kyushu, Shikoku), North Korea; Oriental: China (Fujian, Guangdong, Hainan, Taiwan), Indonesia, Japan (Ryukyu), Malaysia, Sri Lanka, Vietnam; Australian/Oceanian: USA (Hawaii), Micronesia. ( Figs. 319–322 ).

Figures

FIGURES 179 – 183. Homoneura (Homoneura) ornatifrons (Kertész, 1913). Male. 179. body, lateral view; 180, 181. head, anterior and lateral view; 182. thorax, dorsal view; 183. abdomen, dorsal view.

FIGURES 315 – 318. Homoneura (Homoneura) trypetoptera (Hendel, 1908). Male. 315. body, lateral view; 316, 317. head, lateral and anterior view; 318. thorax, dorsal view.

FIGURES 13 – 24. Wing. 13. H. (H.) levis (Wiedemann, 1830); 14. Homoneura (Homoneura) longifurcata sp. nov.; 15. H. (H.) noticomata sp. nov.; 16. H. (H.) pseudograndis Papp & Gaimari, 2013; 17. H. (H.) quadristriata sp. nov.; 18. H. (H.) shunxica sp. nov.; 19. H. (H.) subulifera sp. nov.; 20. H. (H.) tianjingshanica sp. nov.; 21. H. (H.) triangulata sp. nov.; 22. H. (H.) triprocessa sp. nov.; 23. H. (H.) zhangjiajiensis sp. nov.; 24. H. (H.) zhejiangensis sp. nov.

FIGURES 205 – 211. Homoneura (Homoneura) quinquenotata (Meijere, 1915). Male. 205. body, lateral view; 206. head, anterior view; 207, 208. thorax, lateral and dorsal view; 209, 210. abdomen, dorsal and lateral view; 211. wing.

FIGURES 212 – 216. Homoneura (Homoneura) quinquenotata (Meijere, 1915). Male. 212. syntergosternite and epandrium, lateral view; 213. syntergosternite, anterior view; 214. epandrial complex, posterior view; 215. aedeagal complex, ventral view; 216. aedeagal complex, lateral view. Scale bar = 0.1 mm.

FIGURES 1 – 12. Wing. 1. H. (H.) bicuspidata sp. nov.; 2. H. (H.) conspicua Sasakawa, 2001; 3. H. (H.) dadongshanica sp. nov.; 4. H. (H.) degenerata sp. nov.; 5. H. (H.) denticulata sp. nov.; 6. H. (H.) falcata sp. nov.; 7. H. (H.) fengyangshanica sp. nov.; 8. H. (H.) hamata sp. nov.; 9. H. (H.) hongmaoensis sp. nov.; 10. H. (H.) interrupta Sasakawa, 2001; 11. H. (H.) jianlingensis sp. nov.; 12. H. (H.) laticosta (Thomson, 1869).

FIGURES 138 – 142. Homoneura (Homoneura) interrupta Sasakawa, 2001. Male. 138. syntergosternite and epandrium, lateral view; 139. syntergosternite, anterior view; 140. epandrial complex, posterior view; 141. aedeagal complex, ventral view; 142. aedeagal complex, lateral view. Scale bar = 0.1 mm.

FIGURES 217 – 223. Homoneura (Homoneura) quinquevittata (Meijere, 1910). Male. 217, 218. Body, lateral view; 219. head, anterior view; 220. thorax, dorsal view; 221, 222. abdomen, lateral and dorsal view; 223. wing.

FIGURES 272 – 278. Homoneura (Homoneura) strigata (Meijere, 1910). Male. 272. body, lateral view; 273, 274. head, lateral and anterior view; 275, 276. thorax, dorsal and lateral view; 277. abdomen, dorsal view; 278. wing.

FIGURES 279 – 283. Homoneura (Homoneura) strigata (Meijere, 1910). Male. 279. syntergosternite and epandrium, lateral view; 280. syntergosternite, anterior view; 281. epandrial complex, posterior view; 282. aedeagal complex, ventral view; 283. aedeagal complex, lateral view. Scale bar = 0.1 mm.

FIGURES 76 – 80. Homoneura (Homoneura) conspicua Sasakawa, 2001. Male. 76. syntergosternite and epandrium, lateral view; 77. syntergosternite, anterior view; 78. epandrial complex, posterior view; 79. aedeagal complex, ventral view; 80. aedeagal complex, lateral view. Scale bar = 0.1 mm.

FIGURES 97 – 101. Homoneura (Homoneura) degenerata sp. nov. Male. 97. syntergosternite and epandrium, lateral view; 98. syntergosternite, anterior view; 99. epandrial complex, posterior view; 100. aedeagal complex, ventral view; 101. aedeagal complex, lateral view. Scale bar = 0.1 mm.

FIGURES 224 – 229. Homoneura (Homoneura) sauteri Malloch, 1927. Male. 224. body, lateral view; 225. head, anterior view; 226, 227. thorax, lateral and dorsal view; 228. abdomen, dorsal view; 229. wing.

FIGURES 230 – 234. Homoneura (Homoneura) sauteri Malloch, 1927. Male. 230. syntergosternite and epandrium, lateral view; 231. syntergosternite, anterior view; 232. epandrial complex, posterior view; 233. aedeagal complex, ventral view; 234. aedeagal complex, lateral view. Scale bar = 0.1 mm.

FIGURES 235 – 238. Homoneura (Homoneura) septentrionalis (Loew, 1847). Male. 235. body, lateral view; 236. head, anterior view; 237, 238. thorax, dorsal and lateral view.

FIGURES 239 – 243. Homoneura (Homoneura) septentrionalis (Loew, 1847). Male. 239. syntergosternite and epandrium, lateral view; 240. syntergosternite, anterior view; 241. epandrial complex, posterior view; 242. aedeagal complex, ventral view; 243. aedeagal complex, lateral view. Scale bar = 0.1 mm.

FIGURES 260 – 266. Homoneura (Homoneura) simplicissima (Meijere, 1910). Male. 260. body, lateral view; 261, 262. head, anterior and lateral view; 263, 264. thorax, lateral and dorsal view; 265. abdomen, dorsal view; 266. wing.

FIGURES 267 – 271. Homoneura (Homoneura) simplicissima (Meijere, 1910). Male. 267. syntergosternite and epandrium, lateral view; 268. syntergosternite, anterior view; 269. epandrial complex, posterior view; 270. aedeagal complex, ventral view; 271. aedeagal complex, lateral view. Scale bar = 0.1 mm.

FIGURES 71 – 75. Homoneura (Homoneura) concava Sasakawa, 2002. Male. 71. body, lateral view; 72, 73. head, lateral and anterior view; 74. thorax, dorsal view; 75. abdomen, dorsal view.

FIGURES 107 – 111. Homoneura (Homoneura) falcata sp. nov. Male. 107. syntergosternite and epandrium, lateral view; 108. syntergosternite, anterior view; 109. epandrial complex, posterior view; 110. aedeagal complex, ventral view; 111. aedeagal complex, lateral view. Scale bar = 0.1 mm.

FIGURES 112 – 116. Homoneura (Homoneura) fengyangshanica sp. nov. Male. 112. syntergosternite and epandrium, lateral view; 113. syntergosternite, anterior view; 114. epandrial complex, posterior view; 115. aedeagal complex (hypandrium broken), ventral view; 116. aedeagal complex (hypandrium broken), lateral view. Scale bar = 0.1 mm.

FIGURES 133 – 137. Homoneura (Homoneura) hongmaoensis sp. nov. Male. 133. syntergosternite and epandrium, lateral view; 134. syntergosternite, anterior view; 135. epandrial complex, posterior view; 136. aedeagal complex, ventral view; 137. aedeagal complex, lateral view. Scale bar = 0.1 mm.

FIGURES 152 – 157. Homoneura (Homoneura) latifrons Malloch, 1927. Male. 152, 153. body, lateral view; 154. head, anterior view; 155. thorax, dorsal view; 156. abdomen, dorsal view; 157. wing.

FIGURES 289 – 294. Homoneura (Homoneura) subvittata Malloch, 1927. Male. 289, 290. body, lateral view; 291, 292. head, anterior and lateral view; 293. thorax, dorsal view; 294. abdomen, dorsal view.

FIGURES 295 – 299. Homoneura (Homoneura) subvittata Malloch, 1927. Male. 295. syntergosternite and epandrium, lateral view; 296. syntergosternite, anterior view; 297. epandrial complex, posterior view; 298. aedeagal complex, ventral view; 299. aedeagal complex, lateral view. Scale bar = 0.1 mm.

FIGURES 332 – 336. Homoneura (Homoneura) zhejiangensis sp. nov. Male. 332. syntergosternite and epandrium, lateral view; 333. syntergosternite (partly broken), anterior view; 334. epandrial complex, posterior view; 335. aedeagal complex, ventral view; 336. aedeagal complex, lateral view. Scale bar = 0.1 mm.

FIGURES 244 – 248. Homoneura (Homoneura) shunxica sp. nov. Male. 244. syntergosternite and epandrium, lateral view; 245. syntergosternite (partly broken), anterior view; 246. epandrial complex, posterior view; 247. aedeagal complex, ventral view; 248. aedeagal complex, lateral view. Scale bar = 0.1 mm.

FIGURES 190 – 194. Homoneura (Homoneura) pictipennis Czerny, 1932. Male. 190. body, lateral view; 191. head, anterior view; 192. thorax, anterior view; 193. abdomen, dorsal view; 194. wing.

FIGURES 34 – 38. Homoneura (Homoneura) bicuspidata sp. nov. Male. 34. syntergosternite and epandrium, lateral view; 35. syntergosternite, anterior view; 36. epandrial complex, posterior view; 37. aedeagal complex, ventral view; 38. aedeagal complex, lateral view. Scale bar = 0.1 mm.

FIGURES 163 – 167. Homoneura (Homoneura) noticomata sp. nov. Male. 163. syntergosternite and epandrium, lateral view; 164. syntergosternite, anterior view; 165. epandrial complex, posterior view; 166. aedeagal complex, ventral view; 167. aedeagal complex, lateral view. Scale bar = 0.1 mm.

FIGURES 143 – 146. Homoneura (Homoneura) jianlingensis sp. nov. Male. 143. syntergosternite and epandrium, lateral view; 144. epandrial complex, posterior view; 145. aedeagal complex, ventral view; 146. aedeagal complex, lateral view. Scale bar = 0.1 mm.

FIGURES 184 – 189. Homoneura (Homoneura) pallidula Malloch, 1927. Male. 184, 185. body, lateral view; 186, 187. head, anterior and lateral view; 188. thorax, dorsal view; 189. wing.

FIGURES 319 – 322. Homoneura (Homoneura) unguiculata (Kertesz, 1913). Male. 319. syntergosternite and epandrium, lateral view; 320. epandrial complex, posterior view; 321. aedeagal complex, ventral view; 322. aedeagal complex, lateral view. Scale bar = 0.1 mm.