Hlavaciellus decoratus, Jałoszyński, Paweł, 2011

Jałoszyński, Paweł, 2011, Hlavaciellus Jałoszyński, 2006: eleven new species, detailed morphology and systematic position within Cephenniini (Coleoptera, Staphylinidae, Scydmaeninae), Zootaxa 2763, pp. 1-33 : 8-9

Hlavaciellus decoratus sp. n.

( Figs. 10 , 7 , 20, 21 )

Type material. Holotype: MALAYSIA: ɗ, " SABAH \ Mt. Kinabalu \ 1500 m, 21.V.1987 \ Burckhardt - Löbl" [white, printed], " HLAVACIELLUS \ decoratus m. \ det. P. JAŁOSZYŃSKI, '09 \ HOLOTYPUS " [red, printed] ( MHNG).

Diagnosis. Head in males with multiple modifications: vertex separated from frons by pair of broad and deep grooves, each running from posterior margin of eye anteriorly and towards middle and gradually becoming shallower; frons with two large and deep pits; posterior margins of frontal pits separated from median parts of grooves by narrow and recurved carina.

Description. BL 1.83. Body of male ( Fig. 10 ) strongly convex, pigmentation light brown, vestiture yellowish.

Head ( Fig. 7 ) large, HL 0.25, HW 0.48; vertex convex, with highest point clearly defined and located in middle of anterior part, anterior margin broadly V-shaped and separated from frons by pair of deep and broad grooves running from posterior margin of each eye towards middle and anteriorly; anterior margin of grooves separated from frons by narrow and recurved median transverse carina; frons with two large and deep pits; supraantennal tubercles prominent, distinctly separated from frons; eyes large, strongly convex and coarsely faceted. Punctures and setae as in Fig. 7 . Antennae relatively short, about as long as 0.6 BL, slender, with antennomeres VI–XI distinctly broader than I–V, covered with thin, long, dense and suberect setae, additionally antennomeres VII–IX with more erect and straight long setae on internal (e.g. mesal) surface, AnL 1.13, antennomere I 1.4x as long as broad; II distinctly narrower and much shorter than I, slightly broader than long; III–V subequal in length and width, each slightly broader than II and 1.1–1.2 as long as broad; VI distinctly broader and much longer than V, 1.3x as long as broad; VII slightly broader but not longer than VI, nearly as long as broad; VIII slightly larger than VII, as long as broad; IX slightly broader than VIII but not longer, barely noticeably broader than long; X slightly larger than IX, slightly broader than long; XI slightly narrower than X, distinctly shorter than IX–X together.

Pronotum in dorsal view subtrapezoidal with moderately convex anterior margin, broadest at base but barely noticeably narrowing from hind angles to anterior fourth, PL 0.53, PW 0.74; sides strongly rounded in anterior fourth, then nearly straight; hind angles nearly right, blunt; posterior margin shallowly bi-emarginate; ante-basal pits very shallow and diffused, each equally distant from posterior and lateral margin of pronotum; lateral edges barely noticeably swollen. Pronotal disc glossy, in middle covered with very fine and shallow punctures separated by spaces 1– 2 x as wide as puncture diameters, punctures close to margins of pronotum are distinctly denser but not larger and not coarse. Setae long, moderately dense, suberect to erect.

Elytra oval, relatively long, broadest between anterior third and middle, EL 1.05, EW 0.88, EI 1.20; basal pit on each elytron small but distinct, located very close to scutellum; subhumeral lines moderately sharply marked, as long as 0.36x EL, only slightly divergent, each developed as border between higher humeral region and lower adsutural area; apices of elytra separately rounded. Punctures on basal third of each elytron small but more distinct and slightly denser than those on pronotum, gradually reducing in diameter and depth toward lateral margins and apices; setae similar to those on pronotum. Hind wings not studied.

Legs slender and long, all tibiae straight or nearly straight.

Metaventrite with moderately deep postmesocoxal impressions occupying about 2/3 of its length.

Aedeagus ( Figs. 20, 21 ) moderately slender, AeL 0.33; median lobe with abruptly separated, narrow and pointed apical part, in lateral view strongly recurved and strongly narrowing from basal third to apex; internal armature strongly asymmetrical; parameres in lateral view extremely broad, with very slender and strongly curved apical parts.

Female. Unknown.

Distribution. East Malaysia: Borneo, Sabah.

Etymology. The specific epithet decoratus was chosen to reflect the unusually rich suite of secondary sexual characters on the head of the male.

Remarks. The aedeagus and partly the head modifications of this species are similar to those in H. sabahensis . Hlavaciellus decoratus differs clearly in its smaller body length (1.83 vs. 2.10 mm), presence of large frontal pits and large punctures covering the median part of vertex (fine punctures on the frons and vertex in H. sabahensis ).

Figures

FIGURES 10 – 12. Dorsal view of holotypes of Hlavaciellus, males. 10, H. decoratus sp. n. 11, H. smetanai sp. n. 12, H. adelphos sp. n. Scale bars: 1 mm.

FIGURES 4 – 9. Modifications of head in frontal view. 4, Hlavaciellus peninsularis sp. n. 5, H. cirrus sp. n. 6, H. angustifrons sp. n. 7, H. decoratus sp. n. 8, H. smetanai sp. n. 9, H. bifoveatus sp. n. Scale bars: 0.1 mm.

FIGURES 13 – 23. Aedeagus in ventral (13, 15, 18, 20, 22) and lateral (14, 16, 19, 21, 23) views, and head of male in lateral view (17). 13, 14, Hlavaciellus peninsularis sp. n. 15 – 17, H. cirrus sp. n. 18, 19, H. angustifrons sp. n. 20, 21, H. decoratus sp. n. 22, 23, H. smetanai sp. n. Scale bars: 0.2 mm (13 – 16, 18 – 23), 0.1 mm (17).