Hilda transsylvanica (Hoernes & Auinger, 1884)

Harzhauser, Mathias & Landau, Bernard M., 2024, The Colubrariidae, Eosiphonidae, Melongenidae, Pisaniidae, Prodotiidae and Tudiclidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea, Zootaxa 5427 (1), pp. 1-110 : 45-47

publication ID	https://doi.org/ 10.11646/zootaxa.5427.1.1
publication LSID	lsid:zoobank.org:pub:923206B0-E8C5-4FD5-B882-55009ABB0282
persistent identifier	https://treatment.plazi.org/id/03CE9F1C-FFB6-0C69-FF65-FE5BE86EFD4A
treatment provided by	Plazi
scientific name	Hilda transsylvanica (Hoernes & Auinger, 1884)
status

Treatment

Figs 15F View FIGURE 15 , 23A–D View FIGURE 23

* Triton (Hilda) transsylvanicum nov. form.—Hoernes & Auinger 1884: 182, pl. 22, figs 17–20.

Hilda transylvanica [sic] Hoern. et Auing.— Cossmann 1903: 107, plate captions, pl. 5, figs 4–5.

Colubraria transilvanica [sic] Hoernes et Auinger— Korobkov 1955: plate captions, pl. 67, figs 7, 9.

Colubraria (Hilda) transsylvanica (Hoernes et Auinger 1884) —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 139, pl. 38, figs 7a–b.

Pisania transsylvanica (Hoernes & Auinger) View in CoL — Beu & Maxwell 1987: 56, pl. 4, figs j, n, p, q.

Pisania transsylvanica (Hoernes et Auinger, 1884) View in CoL — Kovács 2022: 91, figs 90–93.

Type material. Lectotype (designated herein): NHMW 1876 View Materials /0011/0023a, SL: 22.1 mm, MD: 10.7 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1884: pl. 22, fig. 17), Figs 15F View FIGURE 15 , 23A View FIGURE 23 1 –A View FIGURE 1 3 View FIGURE 3 . Paralectotypes: NHMW 1860 View Materials /0040/0092, SL: 21.1 mm , MD: 10.1 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1884: pl. 22, fig. 18), Figs 23B View FIGURE 23 1 –B View FIGURE 1 2 View FIGURE 2 . NHMW 1865 View Materials /0001/0182, SL: 19.7 mm , MD: 9.6 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1884: pl. 22, fig. 19), Figs 23C View FIGURE 23 1 –C View FIGURE 1 3 View FIGURE 3 . NHMW 1876 View Materials /0011/0023b, SL: 22.8 mm , MD: 10.2 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1884: pl. 22, fig. 20), Figs 23D View FIGURE 23 1 – D View FIGURE 1 2 View FIGURE 2 . Additional paralectotypes: 3 spec. , NHMW 1868 View Materials /0001/0433, Lăpugiu de Sus ( Romania); 21 spec. , NHMW 1870 View Materials /0033/0075, Lăpugiu de Sus ( Romania) .

Revised description. Medium-sized, moderately broad fusiform shell. Apical angle 48–50°. Teleoconch of up to six whorls. Protoconch broad conical of 2.25 convex whorls; diameter: 740 μm, height: 720 μm. Early teleoconch whorls low, with broad, shallow subsutural ramp, angled at shoulder, bearing two prominent spiral cords. Axial ribs widely spaced, most prominent over subsutural ramp and close to abapical suture. Suture deeply incised. Spiral cords swollen at intersections over ribs. Secondary spiral cord appears between primaries on second teleoconch whorl. Additional secondary cord appears on third whorl close to abapical suture. Subsutural ramp with two prominent secondary cords and intercalated tertiary threads. Later whorls convex with periphery below mid-whorl. Primary and secondary cords over subsutural ramp subequal in strength, separated by wide interspaces, with fine tertiary and occasional quaternary threads intercalated. Abapically axial ribs weaken, fading on fifth teleoconch whorl, cords no longer swollen over ribs, resulting in change of sculpture to only spiral cords on last 1.5 whorls. Growth lines strengthen during ontogeny becoming lamellar on last two whorls, forming finely scabrous cancellate pattern. Last whorl moderately high, attaining 65% of total height, evenly convex, moderately constricted at base, bearing 10–13 primary spiral cords with secondary and tertiary threads intercalated; fasciole weak with prominent spiral cords; one broad terminal varix. Aperture ovate, moderately wide. Columella broadly excavated, angled at transition to siphonal canal. Columellar callus forming broad rim, sharply delimited from base in adapical half, with about four weak denticles decreasing in strength adapically; callus thinning in parietal region, not sharply delimited. Anal canal wide U-shape, adjoined by small but prominent parietal denticle and bifid anal denticle. Outer lip thickened with about 13 prominent, elongated denticles starting at lip edge. Adapical eight denticles forming pairs in most specimens; anal denticle most prominent. Siphonal canal moderately short, moderately narrow, slightly bent to the left, moderately notched.

Discussion. Within Paratethyan buccinoids, this species cannot be confused with any other. Dianthiphos sacyi ( Cossmann & Peyrot, 1923) , from the Burdigalian of Saucats ( France) shares the same general shape, although it is slightly slenderer, and the character of the labial denticles is similar, starting at the lip edge and vaguely arranged in pairs. The French species differs most notably in its higher spire and the earlier disappearance of the axial ribs.

Paleoenvironment. Unknown; probably middle to outer neritic settings.

Distribution in Central Paratethys. Badenian (Middle Miocene): Făget Basin: Lăpugiu de Sus ( Romania)

( Kovács 2022); Dacian Basin: Târnene ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).

Genus Janiopsis Rovereto, 1899

Type species. Murex angulosus Brocchi, 1814 , subsequent designation by Sacco (1904: 60). Pliocene, Italy.

Synonyms. Jania Bellardi, 1873 View in CoL [non Jania Lamouroux, 1812 View in CoL , Plantae, but originally described as animal (see Pacaud 2015, 2016].

Revised diagnosis. “ Medium-sized shell (H => 20 mm <60 mm), fusiform. Paucispiral or multispiral protoconch, smooth. Very robust shell, elongated, with convex teleoconch whorls, sculpture formed by axial ribs and distinct spiral cords. Outer lip, internally with folds. Aperture with wide siphonal canal, straight columella, plicate. Umbilicus absent.” ( Brunetti & Della Bella 2016: 27, translated from Italian).

Species included. Fusus maxillosus Bellardi & Michelotti, 1840 View in CoL , Early Miocene, Proto-Mediterranean Sea; Janiopsis vindobonensis nov. sp., Middle Miocene, Central Paratethys Sea; Murex angulosus Brocchi, 1814 , Late Miocene to Pliocene, Mediterranean Sea.

Discussion. Janiopsis is characterized by its peculiar aperture with a large columellar denticle, large parietal and anal denticles, and three very large and blunt denticles in the outer lip. Despite these distinctive features, a very heterogeneous group of species is currently placed in the genus Janiopsis by MolluscaBase Eds. (2021b). Several Paleocene and Eocene species from France have been placed in Janiopsis (as Jania Bellardi, 1873 ) by Pacaud (2015), which differ from the Neogene type by the more numerous and less prominent denticles in the outer lip and the much weaker or obsolete columellar, parietal and anal denticles [e.g., ‘ J. ’ calvimontensis ( Cossmann, 1889) , ‘ J. ’ colridusensis (Pacaud & Leroy in Pacaud, 2015), ‘ J. ’ funiculosa (Deshayes, 1835) , ‘ J. ’ heberti ( Watelet, 1851) , ‘ J. ’ minor ( Deshayes, 1864) , ‘ J. ’ parisiensis ( Deshayes, 1834) , ‘ J. ’ schlumbergeri ( Cossmann, 1889) ]. The numerous denticles in the outer lip distinguish also the Hungarian Eocene ‘ Janiopsis’ dudariensis Strausz, 1966b from Janiopsis . Similarly, the Pliocene ‘ J. ’ hirasei MacNeil, 1961 from Japan and ‘ J. ’ exhexagonus ( Vredenburg, 1925) , from the Early Miocene of Pakistan are clearly not congeneric with the Neogene type species Janiopsis angulosa . Herein we recognize only the Early Miocene Janiopsis maxillosa ( Bellardi & Michelotti, 1840) , the Middle Miocene Janiopsis vindobonensis nov. sp. and the Late Miocene to Pliocene Janiopsis angulosa ( Brocchi, 1814) in Janiopsis . ‘ Janiopsis ’ labrosa ( Bellardi & Michelotti, 1840) , from the Early Miocene of the Colli Torinesi ( Italy), lacks the characteristic columellar denticle and blunt denticles in the outer lip and is excluded herein from Janiopsis ( Bellardi 1873: pl. 11, fig. 7; Ferrero Mortara et al. 1981: pl. 4, fig. 8).