Hemiphileurus

Ratcliffe, Brett C. & Curoe, Daniel J., 2011, A New Species of Hemiphileurus Kolbe (Coleoptera: Scarabaeidae: Dynastinae: Phileurini) from Panama, with a Revised Key to the Panamanian Species, The Coleopterists Bulletin 65 (2), pp. 139-143 : 141-142

KEY TO THE SPECIES OF ADULT HEMIPHILEURUS

OF PANAMA (females of most of the species cannot be identified except by association with the males)

Variation. Male (1 paratype). Length 23.4 mm; width across humeri 11.0 mm. The paratype does not differ significantly from the holotype.

Etymology. This species is named in reference to the country from which it came, but in this case meaning Panamanian rather than “of or from

1. Elytra with alternate intervals strongly elevated, cariniform; 2 rows of large punctures between elevated intervals................ H. dejeani (Bates)

1′. Elytra with all intervals subequal in height; intervals lacking 2 rows of punctures..........2

2. Last sternite moderately to densely punctate......................................................................3

2′. Last sternite punctate or rugose in narrow, transverse band along base only, sparsely punctate elsewhere........................................6

3. Elytra with intervals all similar: straight and smooth...........................................................4

3′. Elytra with intervals dissimilar: occasionally broken, always appearing sinuate because of lateral branching extending to between punctures in furrowed striae..................................5

4. Hollow of frons between tubercles nearly smooth, shiny. Parameres narrow, each with tooth projecting from side ( Figs. 8–9 ) ........... ................... H. pygidiopunctissimus Ratcliffe

4′. Hollow of frons between tubercles punctate, not shiny. Parameres broad, each with large, deep, suboval depression at about middle ( Figs. 2–3 )....... H. panamanius Ratcliffe and ....................................... Curoe, new species

5. Parameres at base each with long, lateral lobe ( Figs. 10–11 )..... H. variolosus (Burmeister)

5′. Parameres at base each lacking a lateral lobe; instead apex of each paramere produced into a lobe ( Figs. 12–13 ) ........... H. vicarius Prell

6. Anterior edge of pronotal furrow with 2 very small tubercles..... H. nebulohylaeus Ratcliffe

6′. Anterior edge of pronotal furrow lacking tubercles....................................................................7

7. Elytra with intervals appearing sinuate because of lateral branching extending between punctures in furrowed striae; elytra appear confusedly sculptured..... H. cylindroides (Bates)

7′. Elytra with intervals similar: nearly straight, smooth, slightly convex................................8

8. Males with horns, not tubercles. Top of head nearly smooth.................... H. simplex (Prell)

8′. Males with tubercles, never with horns. Top of head distinctly punctate to rugopunctate.......9

9. Parameres in caudal view narrowing slightly to apex ( Figs. 6–7 )....... H. laevicauda (Bates)

9′. Parameres in caudal view broadly rounded and bulbous until apex (as in Figs. 4–5 )......10

10. Parameres with elongate, apically expanded, clearly delineated depression at middle of shaft of each paramere ( Figs. 4–5 )............................ ............................................ H. curoei Ratcliffe

10′. Parameres each with declivous area on lateral edges just above middle, not with distinctly edged depression at middle of each shaft ...... .................................... H. dyscritus Ratcliffe

Figures

Figs. 4–13. Caudal and lateral views of the parameres of 4–5) Hemiphileurus curoei; 6–7) Hemiphileurus laevicauda; 8–9) Hemiphileurus pygidiopunctissimus; 10–11) Hemiphileurus variolosus; 12–13) Hemiphileurus vicarius.

Figs. 2–3. Hemiphileurus panamanius, parameres.