Glaresis falli Gordon and Hanley, 2014

Gordon, Robert D. & Hanley, Guy A., 2014, Systematic revision of American Glaresidae (Coleoptera: Scarabaeoidea), Insecta Mundi 2014 (333), pp. 1-91 : 37-38

Glaresis falli Gordon and Hanley , new species

Description. Male. Length 3.4 mm, width 1.8 mm; body form short, slightly elongate, nearly parallelsided, slightly widened from elytral base to apical 1/3 ( Fig. 23A ). Color dark yellowish brown. Head with clypeal surface smooth, dull, frons finely microreticulate, clypeus and frons with pronounced, dense, transversely ovate tubercles, setae short, barely evident; vertex without basal carina, surface coarsely, densely reticulate. Clypeal apex medially emarginate, with large, widely spaced tubercles medially, lateral tubercles small, dense, lateral angles feebly oblique, angulate ( Fig. 23B ). Mandible pair symmetrical; mesal tooth strong; lateral prominence strong, pronounced; outer margin angulate. Pronotum with deep, long, transverse fovea in anterior 1/4 extended completely across pronotum, small slightly transverse fovea on each side of middle in anterior 1/2, wide, deep fovea on each side medially near lateral margin, central furrow deep in basal 1/3, shallow and barely evident in apical 2/3; surface densely rugose, with irregular, mostly vertical, dense, setae-bearing carinae, small, irregularly transverse carinae present in foveae on each side of middle, setae decumbent, about 1/2 length of carinae. Anterior and lateral pronotal margins not bordered, apical and basal margins crenulate. Elytra with surface feebly shiny, densely microreticulate; all striae distinctly, strongly carinate, carinae narrowly interrupted adjacent to space between interval punctures, each carinal segment bearing an apical seta about 1/2 length of segment; intervals with deep, slightly rectangular punctures ( Fig. 23C ). Metasternum long, surface dull, finely, densely microreticulate, not tuberculate medially, laterally with dense, strong, elongate, oblique, setae-bearing ridges, median area flat, with weak carina extended from apical keel 2/3 distance to mesocoxae; metasternal groove weak, distinct, inner margin tapered, outer margin weakly ridged ( Fig. 23D ). Lateral protibial teeth almost evenly spaced, basal two teeth not noticeably close together. Mesotibia with 4 short, widely spaced spines laterally, distinctly projecting at apex, tibial projection apically rounded ( Fig. 23D ). Posterior metatrochanteral margin lacking teeth, posterosuperior metatrochanteral surface with 1 tooth. Metafemoral surface with widely scattered, elongate, setae-bearing tubercles, microreticulate, dull throughout; width to length ratio 1.0:1.6, with narrow flange on anterior margin; posterosuperior margin with 1 very small tooth ( Fig. 23G ). Metatibia broadly triangular, surface entirely microreticulate, outer margin with large, bifid, posteromedian lateral projection, and series of small, regular teeth on lateral margin from base to posteromedian projection, regular row of 5 coarse tubercles extended from base nearly to apex medially, inner margin smooth, pubescent ( Fig. 23F ). Apical margin of 5th abdominal ventrite broadly rounded, barely, almost imperceptibly emarginate medially. Genitalia short, basal piece slightly shorter than parameres, proximal end curved; median lobe slightly shorter than parameres, as wide at middle as paramere, curved upward in apical 1/3, apex acute in ventral view; paramere tapered in lateral view, not flattened, tapered from base to nearly rounded apex ( Fig. 23E ).

Female. Apex of 5th abdominal ventrite with wide, shallow emargination.

Variation. Length 3.0 to 3.4 mm, width 1.7 to 1.9 mm. Size and spacing of apical clypeal tubercles varies slightly from the typical; tubercles on surface of clypeus and frons vary from large, dense, to smaller, more widely spaced; number of tubercles in median row of metatibia vary from 4 to 5.

Type material. Holotype male: California: USA: CALIFORNIA: San Bernardino Co., 0.5mi. E.jct. Riverside Dr ,& Stover St. , Fontana Delhi dune, light, 34 o 03’59"N - 117 o 21’38"W, P.E. Skelley & R. Cunningham ( FSCA) . Paratypes, 74: 8) same data as holotype; (1) USA: CA: San Bernardino Co., Colton Dunes , 34 o 04’N 117 o 18’W, 29 May 2003, A.B.T. Smith; (6) USA:CA: San Berndo , nr. Colton, 34 03’55"N, 117 21’33"W ; sand dunes, May 15. 2010:UV:WBWarner; (35) USA: CA: Riverside Co., Colton Dunes , 312 m, 34 o 03.944’N 117 o 21.957’W, 26 April 2004, Hawks, Ocampo, Paulsen, Smith. (29) USA:CA: San Bernardino Col, Colton Dunes, UV /MV, 26, 29 APR 2004 ; coll. Hawks, Paulsen, Smith, Ocampo; (3) CALI- FORNIA: San Bdo. Co., Colton , dunes Stover Ave, 24 Apr. 2001, D. C. Hawks. ( CMNC) ( FSCA) ( MJPC) ( RCCC) ( USNM) ( WBWC) ( WSCW)

Remarks. This species differs from other group members by a clypeal apex with median tubercles large, widely separated, lateral tubercles small, closely spaced; pronotum with central furrow deep in basal 1/3, weakly impressed in apical 2/3; and metatibial posteromedian lateral projection large, apically bifid. Glaresis falli is present in large numbers at the Colton dunes site which is an unprotected and somewhat damaged set of dunes (R. Cunnngham, pers. com.). The Delhi Sand Dunes were the subject of an article by Longcore (1997), in which he states that they are remnants of the only inland sand due system in the Los Angeles basin, and that the dunes originally extended from Lytle Creek and other small creeks to the Santa Ana River and Jurupa Hills to the south and across the Ontario plain into Loma Linda. Now heavily damaged and reduced in size, the dunes future survival is in doubt.

Etymology. The species is named for H. C. Fall, once a resident of Pasadena, California, who had a single undetermined specimen of this species in his collection (W. Warner, pers. com.). Fall described two species of Glaresis , and was one of the dominant figures in early American coleopterology along with J. LeConte and G. Horn.

Figures

Figures 23A-23G. Glaresis falli, new species. A) Habitus view. B) Clypeus and head surface. C) Elytral striae. D) Metasternum and mesotibia. E) Male genitalia. F) Metaleg, dorsal view. G) Metaleg, ventral view.