Glaresis dakotensis Gordon, 1970

Gordon, Robert D. & Hanley, Guy A., 2014, Systematic revision of American Glaresidae (Coleoptera: Scarabaeoidea), Insecta Mundi 2014 (333), pp. 1-91 : 16-17

Glaresis dakotensis Gordon

Glaresis dakotensis Gordon 1970:503 ; Ratcliffe 1991: 131; Ratcliffe and Paulsen 2008: 64.

Description. Male. Length 3.4 mm, width 2.0; body form robust, slightly widened from elytral base to apical 1/3 ( Fig. 8A ). Color dark yellowish brown. Head shiny, surface punctate from clypeal apex to posterior margin of eye, posterior portion of head finely tuberculate, setae short, barely visible. Clypeal apex sinuate, with narrow median emargination. lateral angles of emargination with low tubercle or slight swelling, entire apex with small, dense, evenly spaced tubercles, appearing serrate, lateral angles oblique, pronounced, outwardly toothed ( Fig. 8B ). Mandible pair symmetrical; mesal tooth strong; lateral prominence strong, pronounced; outer margin abruptly rounded. Pronotum with all foveae weakly impressed except fovea laterally on anterior margin and on each side medially near lateral margin strongly impressed; surface densely rugose, with sparse, seta-bearing carinae, carinae elongate, straight ( Fig. 8C ). Elytra with surface dull, densely microreticulate; striae convex, feebly carinate, carinal segments widely separated, each with short seta; intervals with shallow, slightly rectangular punctures. Metasternum long, feebly shiny, finely, densely microreticulate, surface medially flat without median carina, lateral surface with short ridges in basal 1/2 ( Fig. 8D ); without metasternal groove. Lateral protibial teeth unevenly spaced, basal 2 teeth close together. Mesotibia with 5 spines laterally, 4 spines closely grouped median projection, 1 spine spaced posterior to grouping, tibia strongly projecting at apex ( Fig. 8D ). Posterior metatrochanteral margin without teeth, apical angle without tooth; posterosuperior surface of metatrochanter with single large tooth not visible directly in ventral view ( Fig. 8G ). Metafemoral surface with widely spaced, elongate, seta-bearing tubercles, microreticulate; width to length ratio 1.0:1.4, with narrow flange on anterior margin, large, blunt tooth at angle near trochanter; posterosuperior margin without tooth ( Fig. 8G ). Metatibia broadly triangular, surface entirely microreticulate, outer margin without posteromedian projection, with series of small teeth from base to slightly beyond middle, teeth increasing in size from base with last 3 median teeth largest, apical tooth larger than all others, inner margin smooth, pubescent ( Fig. 8F ). Apex of 5th abdominal ventrite truncate. Genitalia with basal piece slightly longer than parameres, proximal end curved; median lobe nearly as long as paramere, slightly tapered from base to acute apex, wider than paramere at middle, curved upward apically before acute apex; parameres feebly curved in lateral view, apex bluntly rounded ( Fig. 8E ).

Female. Apex of 5th abdominal ventrite feebly emarginate.

Variation. Length 3.4 mm to 4.4 mm, width 2.0 to 2.4 mm. Lateral tubercles on apical clypeal margin often reduced or not apparent, specimens from southern localities tend to have tubercles larger, more distinct than specimens from northern localities; mesotibial spines in posterolateral emargination vary 4-5 in number, and spacing.

Type locality. South Dakota, Hot Springs, Angostura Dam.

Type depository. CNIC.

Temporal distribution. April-August.

Geographical distribution. UNITED STATES. Arizona; Navajo Co., Joseph City ; Colorado: Bent Co., Hasty ; Fremont Co., 4 mi. NE Canon City. Idaho: Power Co., Massacre Rocks State Park . New Mexico: Chaves Co., Bottomless Lake St. Pk ; Eddy Co., 32 o 19.8’N, 103 o 47.3’W ; Eddy Co., Carlsbad ; 26 mi. E. Carlsbad; McKinley Co., Dalton Pass ; mi. N. Tohatchi; Otero Co., Three Rivers Peroglyphs ; Otero Co., 24 km SW Alamogordo ; Otero Co., White Sands ; San Juan Co., Cuaco Canyon ; Union Co., Clayton Lake State Park ; Valencia Co., T7N, R3 W Sect. 30, SE corner . Nebraska: Sioux Co., 17 mi. N. Henry . North Dakota: Slope Co. South Dakota: Fall River Co., Hot Springs, Angostura Dam . Texas: Andrews Co., Andrews ; Brewster Co., Big Bend National Park , Chihuahuan desert nr. Nugent Mt .; Big Bend National Park , Rosillos Lodge ; Big Bend National Park , Tornillo Flat ; El Paso Co., red dunes, 14 mi. NE Fabens ; Presidio Co., Sauceda, Big Bend State Natural Area . Wyoming: Platte Co., Wheatland. ( CMNC) ( CMNH) ( CNIC) ( JCIC) ( MJPC) ( RAAC) ( TAMU) ( UNSM) ( USNM) .

Remarks. This species is most similar to G. phoenicis , see Remarks under that species. Glaresis dakotensis is the eastern version of G. phoenicis , with a distributional range not known to extend farther west than Navajo Co., northern Arizona, and McKinley and Otero counties, New Mexico, then extending north from Texas to North Dakota.

Figures

Figures 8A-8G. Glaresis dakotensis Gordon. A) Habitus view. B) Clypeus and head surface. C) Pronotal surface. D) Metasternum and mesotibia. E) Male genitalia. F) Metaleg and trochanter, dorsal view. G) Metaleg and trochanter, ventral view.