Fastosarion tuljun, Hyman & Köhler, 2019

Hyman, Isabel T. & Köhler, Frank, 2019, Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae), Contributions to Zoology 88 (4), pp. 351-451 : 437-438

Fastosarion tuljun sp. nov.

Figs. 18 , 19O , 32

Etymology

From tuljun (Wiri = hill or mountain), referring to the Tuljunburra (= mountain people) clan of the Wiri people whose traditional lands include the Denham Range; noun in apposition.

Material examined

Holotype. QM MO76081 , Denham Range , ca 2 km NNW of Mt Robert, MEQ. Coll. 18 Dec 2000, D. Cook. G. Monteith.

Diagnosis

External morphology: Shell (fig. 19O) small (8.7–10.2 mm), golden, transparent, ear-shaped, very reduced, 2.6–3.0 whorls, flattened, thin, last whorl large. Body at least 13 mm, discoloured through immersion in pitfall trap; shell lappets and mantle lobes moderately large with some dark markings, smooth. Tail not keeled, slime network moderately strong.

Genital anatomy: Genitalia (fig. 32) with moderately short vagina; bursa copulatrix short, duct distinct, bursa oval. Vagina, free oviduct and base of bursa copulatrix folded into foot. Penis cylindrical, internally with chevron-shaped diagonal ridges distally, a single ridged longitudinal pilaster medially to proximally, proximal portion of penial wall sculptured with transverse ridges; 70% of penis contained in penial tunica. Epiphallus approx. equal to penis length; entering penis through a short, conical verge; epiphallus arms equal in length; epiphallic caecum of moderate length; flagellum of moderate length, slender.

Remarks

This species (identified above as ‘ Helicarionidae sp. Denham Range’) is only known from a single live specimen found in rainforest on the Denham Range, west of Mackay, and several shells from the surrounding region (Mt Bassett, Mt Charlton, Eungella NP) (fig. 18). In the phylogenetic analysis it grouped with F. longimentula and F. sarina . However, morpho-anatomically it is more similar to F. mcdonaldi , both in its small, flattened, reduced shell and in its genitalia (in particular, the presence of a penial verge and the equal length arms of the epipihallus). It can be distinguished from all other Qld semislugs by its unique penis interior.

Figures

FIGURE 18 Distribution of Clade B species. █ = F.alyssa, ▼ = F.deensis, × = F.ephelis, ♢ = F.hannianus, ▲ = F. insularis, ◻ = F.katatonos, ∆ = F.longimentula, ● = F.mcdonaldi, ♠ = F.minerva, ◆ = F.papillosus, ♣ = F.pustulosus, ♥ = F.rowani, Ǫ = F.sarina, ▽ = F.tuljun.

FIGURE 19 Shells of Clade B species. A. Fastosarion alyssa (type locality of D.peterbrocki), NMV F230748. B. Fastosarion deensis, MO85834 (holotype). C. Fastosarion ephelis, MO85835 (holotype). D. Fastosarion hannianus, MO56684.E. Fastosarion insularis sp. nov., MO76354. F. Fastosarion katatonos, MO85837 (holotype). G.Fastosarion longimentula, MO65313.H. Fastosarion mcdonaldi, MO50434.I. Fastosarion minerva, MO78358. J.Fastosarion papillosus, MO78896 (Mt Superbus).K.Fastosarion papillosus, MO62397 (Kroombit Tops). L.Fastosarion pustulosus, MO78894 (Kalpowar SF).M. Fastosarion rowani, MO85839 (holotype). N. Fastosarion sarina, MO85840 (holotype).O. Fastosarion tujun (Denham Range), MO76081 (holotype). Scale bar 5 mm.

FIGURE 32 Genitalia of F.tuljun, MO76081, Denham Range. A. Reproductive system. Arrow shows the junction of the penis and epiphallus.B. Penis interior. Scale bars 1 mm.