Fannia sumapaz, Grisales, Diana, Wolff, Marta & De, Claudio J. B., 2012

Grisales, Diana, Wolff, Marta & De, Claudio J. B., 2012, Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia, Zootaxa 3591, pp. 1-46 : 40-41

publication ID	https://doi.org/ 10.5281/zenodo.213946
DOI	https://doi.org/10.5281/zenodo.6175079
persistent identifier	https://treatment.plazi.org/id/03BE87F8-5216-D401-C094-2287FEE5FAA5
treatment provided by	Plazi
scientific name	Fannia sumapaz
status	sp. nov.

Treatment

Fannia sumapaz View in CoL sp. nov.

( Figs. 35 View FIGURES 20 – 37 , 53 View FIGURES 38 – 55 , 71 View FIGURES 56 – 73 , 89 View FIGURES 83 – 91 , 107 View FIGURES 101 – 109 , 125 View FIGURES 118 – 127 , 130 View FIGURES 128 – 131 )

Diagnosis. these characters apply only to the male sex. Fore tarsus with tarsomeres 4 and 5 flattened and widened, fifth tarsomere larger; hind femur straight, ventral surface without pre-apical protuberance, posterior surface with weak pre-apical protuberance, setulose, with long and weak setae that increase in length on protuberance.

Description. Holotype male: body length 6 mm; wing length 5, 5 mm.

Head: eye densely setulose. Frontal vitta velvety black-brownish pollinose on margins. Interocular space 0.2 mm. Fr 14. Ocellar triangle black; oc proclinate and developed, with 4 less developed setulae on ocellar area. Poc divergent. Fronto-orbital plate and parafacial black,greyish pilose. Parafacial bare and narrowing in ventral third. Fronto-genal suture pronounced and brownish pollinose. Gena brownish pollinose. Lunule black. Scape black with 2 setae reaching to pedicel. Pedicel black with dorsal setae developed. Postpedicel black-brownish pilose, 1.8 times the length of pedicel. Arista black pubescent. Palpus black and claviform.

Thorax: black, weakly greyish pollinose; scutum without vittae and pollinose on postpronotum and scutum. Acr 2:3, not arranged in rows. Dc 2:3 long. 2 Pprn, developed, and 1 smaller. 2 Pra, longer seta near the suture. Prepm setulose. Pre-basal scutellar area with long ground setulae. Subapical scutellar setae 1 pair. Discal scutellar setae 1 pair. Lateral scutellar setae unordered.

Wing: brownish, with upper third between C vein and half of vein R2+3 darker. Calypters whitish with margin of upper calypter brown. Haltere brown with stem base yellowish.

Legs: dark brown with tarsi black and pulvilli brownish. Fore femur with 1 row of long d; 1 row of weak av on basal third; ventral surface bare; 1 row of long pv, uniform and with curved apices; posterior surface setulose, with long and weak setae with curved apices. Fore tibia with 1 pre-apical d; 1 short ad on edge of apical third; 1 short apical v; 1 pre-apical pv; 1 p on edge of apical third. Fore tarsus with 1 strong v on first tarsomere followed by a thin seta; tarsomeres 4 and 5 flattened and widened, fifth tarsomere larger ( Fig. 130 View FIGURES 128 – 131 ). Mid femur constricted on preapical ventral surface; 1 row of ad ending in 3 longer pre-apicals; 1 row of av that decrease in length towards apex, setae on basal third sparse, setae beginning on medium third with hooked apices; 1 row of v on basal third, shorter than the pv; 1 row of long pv with hooked apices, on apical third 2 short and strong setae with hooked apices; 1 row of p long, weak and with curved apices, ending on apical third in 3 longer setae, strong and with straight apices. Mid tibia on basal half of ventral surface constricted, dorsal surface densely setulose on apical half, 1 apical long and strong v; 1 pre-apical developed ad; 1 developed a on edge of apical third and 2 apical a, one long and developed, one short; 1 long pre-apical and strong p; 1 pd on edge of apical third and 1 long and strong pre-apical, same length as apical p. Hind coxa on posterior margin bare. Hind femur straight; ventral surface without preapical protuberance ( Figs. 35 View FIGURES 20 – 37 , 53 View FIGURES 38 – 55 ); posterior surface with weak pre-apical protuberance ( Fig. 53 View FIGURES 38 – 55 ); 1 row of weak a, long and with curved apices, decreasing in length on basal third ( Fig. 35 View FIGURES 20 – 37 ), apical third with long and developed setae going towards dorsal surface; 1 row of weak av with curved apices from base up to median third, apical third with 1 more developed seta ( Fig. 35 View FIGURES 20 – 37 ); ventral surface setulose, with longer setae on basal third; posterior and posteroventral surfaces setulose, with long and weak setae, that increase in length on protuberance ( Fig. 53 View FIGURES 38 – 55 ); 1 row of long pd, weak and with curved apices on basal third, 1 pre-apical. Hind tibia with 1 median long and developed d and 1 shorter pre-apical; 1 median long and developed ad at the same level as d; 1 short pre-apical a; 2 median and 1 apical av ( Fig. 35 View FIGURES 20 – 37 ); 1 short apical v; 1 apical pv, surface setulose from apex down to median third.

Abdomen: elongate, in general with developed setae, black with tergites 3 and 4 basally brown-yellowish, yellowish pollinose mainly on syntergite 1+2. Syntergite 1+2 with a set of long lateral setae; tergites 3 and 4 with 2 pairs of lateral setae, 3 pairs of differentiated median lateral setae; tergite 5 with 1 pair of lateral setae. Sternite 1 densely setulose. Sternite 5 as in Fig. 71 View FIGURES 56 – 73 . Terminalia ( Figs. 89 View FIGURES 83 – 91 , 107 View FIGURES 101 – 109 ): epandrium slightly wider than long, two median sclerotized projections and few setae on basal half; cercal plate round apically, with long and curvilinear setae basally and apically, short lateral setae; surstylus strongly articulated with epandrium, with inner basal projection sclerotized, curved and pointed apically and with weak setae internally; bacilliform process elongate and widened, reaching to the level of surstylus projections; hypandrium and associated structures as in Fig. 125 View FIGURES 118 – 127 .

Female: unknown.

Biology. According to collecting data, Fannia sumapaz sp. nov. is associated with cold ecosystems of the Páramo (up to 3000 m) and occurs at the Parque Nacional Natural Sumapaz . The Páramo ecosystem is characterized by an abundance of water bodies, 40–50 cm grasses, several species of “frailejón ( Asteraceae , Espeletia ) and a high diversity of plant and animal species (Parques Nacionales Naturales 2002).

Comments. These characters apply only to the male sex. Fannia sumapaz sp. nov. is close to F. i g u a q u e sp. nov., but differs by the smooth ventral surface of the hind femur, without a protuberance, and by the absence of a tuft of developed setae. The terminalia of Fannia sumapaz has inner basal projections on the surstylus, a character shared with the species of the hirticeps group, which have a Nearctic and Palearctic distribution, and which are close to the lugubrina group ( Chillcott 1961; Rozkošný et al. 1997). However, the monophyly of this group has not been tested and, according to Chillcott (1961), the hirticeps group is not homogenous in composition.

Etymology. Name in apposition. The species epithet refers to the name of the Parque Nacional Natural Sumapaz , one of the main ecosystems of the Páramo of Colombia, and the type-locality of the species. The park is located between the departments of Cundinamarca, Meta and Huila, at the Central Cordillera.

Type material. Holotype male (IAvH).CO [ Colombia]. Cundinamarca. P.N.N [Parque Nacional Natural] Sumapaz \ Bocatoma. Cerro El Zapato\ 4˚14’N 74˚ 12W. 3560 m. Pantra. 6–7.XI.2002 \H. Angel y D. Arias. Leg. M. 798. IAvH. Paratype: CO [ Colombia]. Cundinamarca. P.N.N [Parque Nacional Natural] Sumapaz . Bocatoma\Cerro El Zapato. 4˚14’N 74˚12W\ 3560 m. Malaise. 2–17.i.2003 \A. Patiño\ Leg. M. 3442. (1 male, IAvH). The holotype is not in a good state, but has all structures.

Distribution. Colombia: departments of Cundinamarca (Andean region, Eastern Cordillera) and Meta (Amazon region).