Exomala (Neoblitopertha) campestris ( LATREILLE , 1804)

Rössner, Eckehard, Zorn, Carsten & Branco, Tristao, 2009, Exomala (Neoblitopertha) campestris (L, 1804) and Exomala (Neoblitopertha) succincta (C, 1840): two distinct European species (Coleoptera, Scarabaeidae, Rutelinae, Anomalini), Beiträge Zur Entomologie = Contributions to Entomology 59 (1), pp. 175-189 : 178-180

Exomala (Neoblitopertha) campestris ( LATREILLE, 1804)

Melolontha campestris LATREILLE, 1804: 195 (type locality: „midi de la France “).

Anisoplia arenaria CASTELNAU, 1840: 151 (type locality: „ France méridionale“).

Phyllopertha campestris var. occidentalis MULSANT, 1842: 497 (type locality: Bordeaux and Mont-de- Marsan).

Phyllopertha campestris var. circumcincta MULSANT, 1842: 497 (type locality: Bordeaux and Mont-de- Marsan).

Phyllopertha campestris var. sabulosa MULSANT, 1842: 497 (type locality: Bordeaux and Mont-de-Marsan).

Blitopertha campestris var. hispanica BÁGUENA, 1955: 293 (type locality: Spain); LEPLAT, 1958: 60 [ Blitopertha campestris hispanica ].

Blitopertha campestris var. mulsanti BÁGUENA, 1955: 293 (type locality: Bordeaux and Mont-de-Marsan).

Examined material:

France: „Gall. mer., occ. ...[partly illegible] | 24645“ (1 , historical collection, MNHB). – „24645“ (2 , historical collection, MNHB). – „Frankreich“ (1 , 2 , MNHB). – „Gall. austr.“ (1 , coll. Hänel, SMTD). – „ Gironde: Cap Ferret, dune océan, cote 99, fleurs, 15.VI.[19]66“ (1 , CTB). – „ Gironde: La Teste, 4.VII.[18]84“ (1 , CTB). – „ Gironde: Le Haillan, 20.VI.[19]43, G.Tempère legit“ (1 , CTB). – „ Gironde: Le Pilat, VI.[19]47, G.Tempère legit“ (1 , CTB). – „ Gironde: Le Porge, 11.VI.[19]61, J. Aubry legit“ (2 , CTB). – „ Gironde: St. S. de Cadourne, à Barclès en fouettant, VI.[19]24“ (2, CTB). – „ Gironde: Soulac-sur-Mer, H.Gouin legit“ (1 , CTB). – „ Gironde: Villeneuve d’Ornon, 2.VII.1926, G.Tempère legit“ (1 , CTB). – „Haute-Garonne, 1924, Duffort legit (ex col. G.Tempère)“ (1 , CTB). – „ Landes, J. Feytaud legit“ (2 , CTB). – „Landes: env. de Mont de Marsan , 1955, J. Feytaud legit“ (2 , 1 , CTB). – „Cauterets, Hautes Pyrenées | Coll. Piesbergen “ (1 , 1 , SMNS). – „Süd Europa | Coll. Jäger “ (1 , 3 , SMNS) .

Andorra: „ Andorra e.V. 1.7.[19]10 J | Sammlung J. Daniel“ (4 , 9 , ZSR).

Spain: „ Spanien , Prov. Lerida : Sepeira (12 km nw Tremp), 1200 m, 22.VI.1992, leg. H. Kalz “ (3 , CHK) . – dto., nur „leg. A. Schröder “ (1 , CER). – „ Spanien , Prov. Teruel : Mas de las Matas (13 km sö Alcorisa), 500 m, 17.-21.VI.1992, leg. E. Rössner “ (1 , 1 , CER). – „Ponferrada, Paganetti“ (1 , SMTD) . – „ Hispania “ (1 , coll. Felsche, SMTD) .

Description:

Head, pronotum and scutellum black, shining, often with weak metallic lustre. Elytra yellowish brown, moderately shining; dark marks usually weakly developed: narrowly darkened along the periphery, umbone with a dark mark, the dark transversal band sometimes indicated.

Clypeus broadly rectangular; sides parallel; fore edge reflexed and weakly convex; anterior angles rounded; punctation transversely rugose.

Forehead and vertex with erect, light-coloured pilosity; with very dense, rugose punctation. Antennal club elongate; outer face of last antennomere weakly convex.

Pronotum ( fig. 5 ) evenly rounded at the sides; base convex, completely margined, margin extending to the postero-lateral angles though here sometimes very thin; punctation transversely dense to rugose.

Scutellum ( fig. 5 ) most often elongate, sometimes semicircular; densely punctate, setose.

Elytra ( figs 11-16 ) broadest approximately in the middle; odd intervals feebly convex and weakly setose; all intervals with more or less distinct transverse, rugose punctation.

Pygidium densely transversely rugose; with moderately dense, long light-coloured pilosity.

Ventral face and femora with rather long light-coloured setae.

Inner claw of fore tarsi in males ( fig. 4 ) relatively short, less than half as long as last tarsal joint.

Figures

Figs 1-5: Exomala campestris, France, Gironde, La Teste, . – 1: Aedeagus, dorsal. – 2: Aedeagus, ventral. – 3: Aedeagus, lateral. – 4: Last joint and inner claw (outer claw not shown) of right fore tarsus. – 5: Pronotum and scutellum.

Figs 11-22: Colour pattern of left elytron. – 11-16: Exomala campestris – 17-22: Exomala succincta – 11-13, 17-19: . – 14-16, 20-22: . – 11, 14, 17, 20: Specimens with reduced dark marks. – 12, 15, 18, 21: Specimens with moderately developed dark marks. – 13, 16, 19, 22: Specimens with well developed dark marks. – 11: Gallia mer. – 12: France. – 13: Gironde: La Teste. – 14: France. – 15: France. – 16:Wironde: Soulac-sur-Mer.– 17: Tirol. – 18:Tirol mer. – 19: Tirol. – 20: Emilia, Reno. – 21: Piemonte: Garearel. – 22: Bolzano.