Exocelina fume (Balke, 1998)

Shaverdo, Helena, Sagata, Katayo & Balke, Michael, 2018, Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae), ZooKeys 803, pp. 7-70 : 19

7. Exocelina fume (Balke, 1998) Figs 3, 27

Copelatus (Papuadytes) fume Balke, 1998: 330; Nilsson 2001: 76 (catalogue).

Papuadytes fume (Balke, 1998): Nilsson and Fery 2006: 56 (comb. n.).

Exocelina fume (Balke, 1998): Nilsson 2007: 33 (comb. n.); Nilsson and Hájek 2018: 66 (catalogue).

Type locality.

Papua: Pegunungan Bintang Regency, Borme, 04°24'S, 140°25'E, 1800 m a.s.l.

Type material studied.

Holotype: male "IRIAN JAYA Zentralmassive 140°25'E 04°24'S, "Borme, 1800m 16.8.1992 leg. Balke (12, 12A)", “HOLOTYPUS” [red], " Copelatus fume Balke des. 1997" [red] (NHMW). Paratypes: 9 males with the same label as the holotype and additionally with red labels "Paratypus Copelatus fume Balke des. 1997", two of them with labels "M.Balke 3275" [green], and "M.Balke 3276" [green] and "M.Balke 6405" [green text] (NHMW). Note: There are two additional paratypes of E. fume , which do not belong to this species but to E. ketembang (Balke, 1998) and E. erteldi (Balke, 1998).

Diagnosis.

For complete description, see Balke (1998: 330). Beetle medium-sized: TL-H 3.7-4.4 mm; oblong-oval; brown to dark brown, with reddish brown pronotal sides, head, and sometimes also sides of elytra, in some specimens, disc of pronotum and elytron almost piceous; submatt, with fine but rather dense punctation and strongly impressed microreticulation; pronotum without lateral bead; male antennae simple (Fig. 3); male protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta; male protarsomere 5 long and narrow, with anterior band of more than 60 and posterior row of 6 relatively long, thin setae (Fig. 27D); median lobe in lateral view slightly curved, with apex curved downwards, with visible angle on dorsal side, in ventral view, distally distinctly narrowed to truncate apex; paramere slightly concave on dorsal side and with long, dense, thin setae, situated along dorsal margin: subdistal setae strong and dense, setae in middle part shorter and sparser, proximal setae long, only slightly sparser than subdistal ones (Fig. 27 A–C).

Affinities.

In the area of its distribution, E. fume co-occurs with E. takime (Balke, 1998) and species of the ekari -, aipo-, okbapensis-, aipomek-, erteldi -, and danae -groups. From species of the ekari -group, the species differs in larger size, evidently stronger dorsal punctation and microreticulation, and the shape of the median lobe. In the latter two characters, E. fume differs also from the species of the remaining groups, as well as in absence of the pronotal bead and simple male antennae.

Within the casuarina -group, the species is more similar to E. casuarina and E. ibalimi sp. n., especially the latter one, from which can be distinguished by paler coloration and the shape of the median lobe (see their “Affinities” and the “Key”).

Distribution.

Papua: Pegunungan Bintang. The species is known only from the type locality (Fig. 50).