Euspinoliina Brothers & Lelej, 2017

Brothers, Denis J. & Lelej, Arkady S., 2017, Phylogeny and higher classification of Mutillidae (Hymenoptera) based on morphological reanalyses, Journal of Hymenoptera Research 60, pp. 1-97 : 11

Euspinoliina Brothers & Lelej subtrib. n.

Type genus.

Euspinolia Ashmead, 1903. This group was found to be monophyletic in all analyses with very high resampling support (here GC = 90), but somewhat inconsistent in its placement; our justification for including it in the Pseudomethocini appears above. The close association of Atillum and Hoplocrates has long been recognized, but the inclusion of Euspinolia with them and separate from the other pseudomethocines is unexpected. The group is not supported by any unique and unambiguously placed synapomorphies, but there is one unique but ambiguously placed synapomorphy for both additive and non-additive characters: 163.2, fore tibia with obliquely elongate outer secretory pore in males (but absent in some Euspinolia ). There are 13 unambiguously placed homoplasious synapomorphies, the most significant being: 70.3, fore tibia with obliquely elongate outer secretory pore in females (also only in Ronisia ); 82.0 and 201.0, tergum I and/or propodeum with simple pubescence in females and males (within Sphaeropthalminae also only in Cephalomutilla , Gogoltilla , some Dasymutilla , some Bothriomutilla females, and Lophomutilla males); 99.0, head with simple pubescence in males (within Sphaeropthalminae also only in Cephalomutilla ); 219.1 and 220.1, gonostylus (paramere) short, tapered and apically straight (within Sphaeropthalminae also only in Myrmilloides ). The subtribe is Neotropical, with three genera; females and males are known for all genera.