Eudorylas skorpionensis, FÖLDVÁRI, MIHÁLY, 2013

FÖLDVÁRI, MIHÁLY, 2013, Taxonomic revision of the Afrotropical species of the tribe Eudorylini (Diptera, Pipunculidae), Zootaxa 3656 (1), pp. 1-121 : 101-102

Eudorylas skorpionensis sp. nov.

( Figs 35A–F )

Diagnosis : One pale anteromedial hair on 3rd tibia; S7 visible. Eyes touching for distance equal to 1–1.5 times ocellar triangle. The highly asymmetric gonopods are characteristic for this species, IG has a long process curved towards the median line (always with a flap-like structure at the base) and the PG is broad, S-shaped in lateral view. Type material: Namibia: 1♂, HT [# T647], Lüderitz, Skorpion Hill , 27°49’S 16°36’E, 09–12.viii.1997., Marais & Kirk-Spriggs, Malais trap. ( NMNW) ; 2♂, PTs, same data as HT ( NMNW, HNHM) .

Other material examined: Namibia: 1♂, S.W. Africa (10), Plateau Fm., 22 mls. E.Aus, 14–17.i.1972; Southern African Exp. B.M. 1972-1. (two labels by Hardy with notes and identification as Pipunculus (Eudorylas) sp.n. nr. amuscarium) ( BMNH); 1♂, same collecting data, without Hardy's labels ( HNHM); 1♂, 2218Ca, Windhoek Dist. Windhoek, 1600m, 3-11-1974, ME Irwin, sandy river bottom; 1♂, 2617Cd, Bethanien Dist. 45km. W. Seeheim, 800m, 19-II-1974, ME Irwin, sandy river bank (both NMSA) ; 6♂, 2617 Ca, Bethanien Dist. 15km. W. Goageb, 1100m, 19-II-1974, ME Irwin, sandy river bank (4 NMSA, 2 HNHM) . South Africa: 2♂, Cape Prov 14mi. N. Springbok, 2450ft, Sept.6, 1972, 2917 Bd, M.E. Irwin, open hillside (both NMSA) ; 3♂, Cape Prov , 11mi. NNE Hondeklipbaai, Sept.8, 1972, 3017 Ab, ME &BJ. Irwin, 200ft. alt-, Reddish sand, shrubs (2 NMSA, 1 HNHM) .

Male

Head. Third antennal segment acuminate; brown, more yellowish towards tip. Face silvery pollinose. Frons, upper part shining black, lower part silvery pollinose; eyes touching for distance equal to 1–1.5 times ocellar triangle. Occiput, lower half silvery pollinose, upper half subshining black.

Thorax. Humeri pale brown. Mesonotum (viewed obliquely from front) subshining black, along anterior margin greyish pollinose; completely silvery from the side. Scutellum subshining black, with 4–5 pairs of short, pale hairs. Dorsocentral hairs weakly developed, notopleuron with several whitish hairs (as long as t3 at base). Halter yellowbrown, stem darker.

Legs. Trochanters and femora brown (all femora shining ventrally, f3 also posteriorly), distal 1/5 of femora and knees yellow, tibiae and tarsal segments yellow-brown, last segment brown. Ventroapical row of 6–7 very short, black spines on mid femora; no spines on 1st femora, only white some hairs on 3rd femur. Subapical (distal) spines on first four tibiae present. One pale anteromedial hair on 3rd tibia. Hind trochanter with several pale hairs on shining brown ventral side. Pulvilli as long as than last tarsal segment.

Wing. Fourth costal section 0.6 times as long as third costal section. Cross-vein R-M just beyond 1/4 of discal cell. Pterostigma coloured on distal 4/5. Hairs on tegula missing.

Abdomen. Viewed obliquely from front tergites brownish pollinose (except T1 being grey), T1–4 greyish pollinose on distal 1/ 4 in the middle, distal 1/2 laterally; distal half of T5 completely grey, sides silvery pollinose. Hairs dispersed, short and weakly developed. Laterally 2–3 brown spines on first tergite (as long as 2 times the width of f3 at base). Postabdomen in dorsal view: S7 visible; T5 as long as ST8. Genitalia without dissection: ST8 silvery pollinose, roundish, but angled on right side dorsally and with a small depression on the right proximal edge, no m.a., SS and EP yellow.

Genitalia. Surstyli short, asymmetrical; epandrium enlarged; ST8 narrow, without m.a. ( Fig. 35E ). SES developed, without distinct hairs; gonopods highly asymmetrical, IG with a long process bent medially (the base of the process bears always a flap-like structure), OG very small, rounded; hypandrium with elongated lobes ( Fig. 35F ); phallic guide broad, narrowing towards tip, S-shaped in lateral view ( Fig. 35A ); phallus supported by membranous structures, trifid, strongly curved, but not coiled distally; ejaculatory apodeme triangle shaped; sperm pump oval, with two lateral projections ( Fig. 35D ).

Female—Unknown.

Remarks —The PG of this species may be interpreted as the OG, since it is lateral in position and no other structure is present between the two gonopods. This feature is also present in the closely related E. bipertitus Kehlmaier, 2005 from Israel.

Distribution— Namibia, South Africa.

Etymology —The species was named after the hill where the type specimens have been caught.

Figures

FIGURE 35. Male genitalia of Eudorylas skorpionensis sp. nov. (A–F) and E. swanengi (G–N).—A: outer surstylus with phallic guide, phallus and outer gonopod, lateral view, B: surstyli, dorsal view, C: inner surstylus, lateral view, D: sperm pump with ejaculatory apodeme, E: dorsal view, F: ventral view with gonopods (without ST8), G: outer surstylus with phallic guide and outer gonopod, lateral view, H: ventral view with phallic guide and subepandrial sclerite (without ST8), I: inner surstylus, lateral view, J: sperm pump with ejaculatory apodeme, K: surstyli, dorsal view, L: dorsal view, M: phallic guide, ventral view, N: phallus, lateral view. Scale bars: 0.1 mm