Eryma sulcatum Harbort, 1905

Devillez, Julien, Charbonnier, Sylvain, Hyžný, Matúš & Leroy, Lucien, 2016, Review of the Early Cretaceous erymid lobsters (Crustacea: Decapoda) from the Western Tethys, Geodiversitas 38 (4), pp. 515-541 : 521-522

Eryma sulcatum Harbort, 1905

( Fig. 4 I-L)

Eryma sulcata Harbort, 1905: 15 , pl. 1, fig. 11, pl. 11, fig. 4. — Glaessner 1929: 159. — Woods 1930: 80, pl. 22, figs 5-7. — Van Straelen 1936: 9. — Förster 1966: 124, fig. 23, pl. 17, figs 2, 4. — Taylor 1979: 34. — Feldmann & Titus 2006: 64.

Astacodes falcifer (pars) – Bell 1863: pl. 9, figs 7, 8.

Eryma cf. sulcata – Aguirre-Urreta & Ramos 1981: 610, pl. 1, fig. a.

Eryma sulcatum – Schweitzer et al. 2010: 25. — Karasawa et al. 2013: 102.

TYPE MATERIAL. — Neotype herein designated SM B11437, specimen figured by Woods (1930: pl. 22, fig. 5a, b). After Harbort (1905), the original type material was composed of at least three specimens from the Hauterivian of Stadthagen near Schaumburg (Lower Saxony, Germany). AfterFörster (1966: 124) and Morris (1980: 7), these specimens were destroyed during World War II and only poorly preserved casts are housed at the NHM, London (see Woods 1930: 80 and Förster 1966: 124). Förster (1966) designated a lectotype based on a cast corresponding to the carapace figured by Harbort (1905: pl. 1, fig. 11a, b). After examination, the quality of this cast is not good enough to be used and we consider important to select a neotype for Eryma sulcatum . We select the historical specimen SM B11437, which is three-dimensionally preserved and fits the original diagnosis according toWoods (1930), Förster (1966) and to our own observations ( Fig. 4I, K ). The new type locality is Speeton in northern England. After Woods (1930), the specimen was collected in the C4 bed of the Speeton Clay, which corresponds to the Hauterivian after Fletcher (1969).

TYPE LOCALITY. — Speeton, Yorkshire, United Kingdom.

TYPE AGE. — Hauterivian, Early Cretaceous (Speeton Clay C Beds).

DESCRIPTION

Subcylindrical carapace (neotype: CL = c. 32 mm, visible CH = c. 12 mm); short rostrum; fusiform, tuberculate intercalated plate limited by a row of coarse tubercles; almost smooth post-orbital area; cephalic region with oblique orbital row of tubercles ending by strong orbital spine (antennal row absent); deep cervical groove, intercepting dorsal margin at c. 66° angle, joined to antennal groove; short gastro-orbital groove originating as a slight median inflexion of cervical groove; postcervical groove slightly convex forward, interrupted before joining dorsal margin, joined medially to branchiocardiac groove; deep branchiocardiac groove, strongly inclined, interrupted before joining dorsal margin, joined to hepatic groove; concavo-convex hepatic groove; slightly inflated ω bulge, delimited ventrally by a narrow and shallow depression extending between antennal and hepatic grooves; slightly inflated χ bulge; inferior groove curved forward; carapace showing dense ornamentation of small tubercles and pits; branchial region with finer tubercles; pleon densely covered with small numerous tubercles and pits; s1-s3 pleura with pointed ventral margin; pleonal somites with terga densely covered by small pits; longitudinal bulge above pleura basis; subtriangular pleura with pointed ventral margin, wide and short in s2, narrower and longer in s3 and s4, densely covered with small pits; chelate P1; massive P1 propodus, longer than wide, relatively thick with rounded inner and outer margins; P1 palm densely covered with small tubercles and deep pits; fingers not preserved; short P1 carpus, showing coarser ornamentation than propodus, distal extremity of external margin bearing a strong spine; P1 merus with the same ornamentation than P1 carpus, with spines on the dorsal margin and a spine on the distal extremity of external margin; slender, P2-P5 with a small punctation.

DISCUSSION

Eryma sulcatum shows a heterogeneous ornamentation (branchial region with fine tubercles, cardiac and cephalic regions with coarser tubercles). This observation is not common among most of the representatives of Eryma , which exhibits more homogeneous and fine ornamentation on the whole carapace. It is worth mentioning that several Early Cretaceous species of Eryma exhibit the same heterogeneous and/or coarse ornamentation such as E. glaessneri (cephalic and cardiac regions with dorsal rows of strong tubercles; branchial region with fine tubercles) or E. nippon (coarse ornamentation).

Figures

FIG. 4. — Species of Eryma Meyer,1840 from Western Europe.A-D, holotype MNHN.F.A57457 (Clément coll.) of Eryma vocontii n. sp.from the Albian of Rosans,France: right lateral view (A), line drawing (B), left lateral (C) and dorsal (D) views; E-F, paratype MNHN.F.A57458 (Clément coll.) of Eryma vocontii n. sp.: dorsal view of right P1 chela (E) and line drawing (F); G, H, cast of the holotype MNHN.F.R10204 of Eryma glaessneri (Van Straelen,1936) from the Hauterivian of Escragnolles,France:left lateral view (G) and line drawing (H); I-L, neotype herein designated SM B11437 of Eryma sulcatum Harbort, 1905 from the Hauterivian of Speeton, United Kingdom: left lateral view (I), dorsal view (K) and line drawings (J, L). Abbreviations:a, branchiocardiac groove; b, antennal groove; b1, hepatic groove; c, postcervical groove; d, gastro-orbital groove; e1e, cervical groove; i, inferior groove; ip, intercalated plate; PoA, post-orbital area; ω, attachment site of mandibular muscle; χ, attachment site of adductor testis muscle. Photographs:L. Cazes (A-D,G), J. Devillez (I, K). Line drawings:J. Devillez. Scale bars: A-F,5 mm; G-L, 10 mm