Epichrysomallidae new status, 1964

Burks, Roger, Mitroiu, Mircea-Dan, Fusu, Lucian, Heraty, John M., Jansta, Petr, Heydon, Steve, Papilloud, Natalie Dale-Skey, Peters, Ralph S., Tselikh, Ekaterina V., Woolley, James B., van Noort, Simon, Baur, Hannes, Cruaud, Astrid, Darling, Christopher, Haas, Michael, Hanson, Paul, Krogmann, Lars & Rasplus, Jean-Yves, 2022, From hell's heart I stab at thee! A determined approach towards a monophyletic Pteromalidae and reclassification of Chalcidoidea (Hymenoptera), Journal of Hymenoptera Research 94, pp. 13-88 : 13

Epichrysomallidae new status

Epichrysomallinae Hill & Riek, 1967. Type genus: Epichrysomalla Girault, 1915.

Diagnosis.

Antenna with 10-12 flagellomeres, including a small 4th clavomere. Eyes not ventrally divergent. Labrum hidden behind clypeus, flexible. Mandibles with 3 teeth. Subforaminal bridge with postgenal bridge separating secondary posterior tentorial pits from hypostoma. Notauli complete. Mesoscutellum with frenum indicated laterally, without axillular sulcus (Fig. 17 ). Mesopleural area without an expanded acropleuron; mesepimeron extending over anterior margin of metapleuron (Fig. 17 ). All legs with 5 tarsomeres in most, except tarsi 4-segmented in Odontofroggatia Ishii and Josephiella Narendran; protibial spur stout and curved; basitarsal comb longitudinal. Metasoma with syntergum, therefore without epipygium.

Discussion.

Epichrysomallidae mostly resemble Melanosomellidae in habitus, but do not have a linear mesopleural sulcus. They also have different fore wing venation with a stigmal vein arising at a right angle (excepted in Acophila Ishii) and a postmarginal vein that is shorter than the stigmal vein. Epichrysomallidae have a characteristic flap-like expansion of cuticle from the lateral edge of the propodeal spiracle, partially covering the spiracle in dorsal view (Fig. 18 ) that neither Melanosomellidae nor Ormyridae have. Ormyridae differ further from Epichrysomallidae by having a more conventional fore wing venation, with longer marginal and postmarginal veins, and iridescent coloration in most species.

Figures

Figures 13 - 18. 13 Coelocyboides sp. (Coelocybidae): mesosoma in dorsal view, arrow indicating setae on or adjacent to frenal groove 14 Ormyromorpha trifasciata Girault (Coelocybidae): mesosoma in lateral view 15, 16 Lelaps sp. (Diparidae) 15 head posterior view 16 cercal brush 17 Eufroggattisca polita (Ashmead) (Epichrysomallidae): mesosoma lateral view 18 Odontofroggattia sp. (Epichrysomallidae): propodeum female dorsal view.