Eoclivina schmidli, Balkenohl, 2023

Balkenohl, Michael, 2023, The genus Eoclivina (KULT, 1959) from Africa with the description of two new species (Coleoptera, Carabidae, Clivinini), Belgian Journal of Entomology 138, pp. 1-17 : 13-15

Eoclivina schmidli sp. nov.

( Figs 5 , 9 , 14 )

urn:lsid:zoobank.org:act:56BA6969-1ADF-49E4-9281-16DCFEBB20DB

ETYMOLOGY The name is a patronym of the German entomologist Dr. Jürgen Schmidl to recognize his support and technical advice of my work over the last 30 years as well as supporting me with numerous material from his excursions.

TYPE MATERIAL. Holotype: ♂, with white black framed and printed and handwritten label and data: “Tanganika D.O.AFRIKA Böttcher” / “Sammlung H. HESSE SMNS 1995 ” / “ Clivina dumolinii-group spec. det. Balkenohl, 2001 ” ( Fig. 5 ) ( SMNS).

Paratypes: 2 ♀♀, same data as holotype but “ Iringa ” ( SMNS, CMBB).

REMARK. In the paratypes, some tarsalia are missing at the right intermediate and hind leg.

DIAGNOSIS. A large sized piceous species with unicolor elytron and interval eight of the elytron reticulated. It is distinctly the largest sized species of the genus and is distinguished besides the size from the most similar species E. machadoi and E. burgeoni by the different reticulation pattern on the elytron, the glabrous disk of the pronotum, and the different male aedeagus and the female gonocoxite.

DESCRIPTION. Measurements. Holotype: Body length 8.43 mm, width 2.46 mm; ratio L/W of pronotum 1.0; ratio L/W of elytra 1.85. Paratypes: Body length 8.32 mm, 8.69 mm; width 2.56 mm, 2.44 mm; ratio L/W of pronotum 1.0, 1.0; ratio L/W of elytra 1.87, 1.78.

Colour: Glossy. Piceous, antenna, mouthparts, intermediate and hind leg fuscous. Supraantennal plate translucent-fuscous.

Head: A third narrower than pronotum. Clypeus with central part bilobed anteriorly; lateral lobe distinctly projecting, wing indistinct, almost fused with central part; separated from supraantennal plate by right-angulated notch, supraantennal plate laterally regularly convex, laterally less projecting than eye; clypeus, wing, and supraantennal plate reflexed margined. Supraantennal plate overlapping eye latero-anteriorly by a quarter (dorsal view), extended posteriorly into a keel, separated from wing by a flat furrow. Clypeus transverse, convex, separated from frons by indistinct flat carina in form of an inverted V, followed posteriorly by a furrow. Frons convex, with a triangle-like impressions at middle, supraantennal plate continuing uninterrupted as widened supraorbital carina up to posterior eye-level. Clypeus smooth, frons with few micro-punctures, separated from supraantennal plate and supraorbital carina by deep and wide furrow, with clypeal seta in front; with two supraorbital setigerous punctures at middle and posterior end of the eye. Neck constriction missing. Eye distinctly convex in dorsal view. Main part of eye situated ventrally where it is globose. Due to the overlapping supraantennal plate, the eye resembles in lateral view anteriorly contracted. Gena distinct, convex, covering a quarter of posterior eye in ventral view. Antenna short, not reaching middle of pronotum. Labrum nearly straight but slightly advanced at middle, with transverse to irregular reticulation, six setose. Mentum with lateral lobe asymmetric, base isodiametrically reticulated, at middle with transverse reticulation, lateral lobe flattened, with longitudinal to irregular reticulation. Median tooth shorter than lateral lobe, wide and hollowed out at apex, with long carina at base.

Pronotum: Disk convex in lateral view, distinctly convex in frontal view with slightly increasing convexity laterally. As long as wide, peltate. Reflexed lateral margin smooth, somewhat attenuating in anterior half with slight convexity, widest behind middle; anterior angle distinct, rounded and slightly projecting, posterior angle missing; lateral channel narrow, completely and regularly rounded off at posterior setigerous puncture and continuing up to base. Median line narrow, sharp, complete. Anterior transverse line consisting of longitudinal connected punctures, joining median line, not joining anterior margin. Surface smooth, with few micro punctures, without punctures in basal half, with a few transverse wrinkles. Base narrow but distinct, three times as wide as lateral channel. Proepisternum just not traceable in dorsal view.

Elytron: Disk in lateral view flattened in anterior half and regularly convex in posterior half, distinctly and regularly convex in frontal view. Shape long-oval with indistinctly convex and diverging part in anterior half, with maximum width slightly behind middle. Humerus rounded, distinct. Reflexed lateral margin smooth. The row of setigerous punctures in the lateral channel is interrupted at middle by one puncture. Scutellar striole just traceable at declivity; setigerous tubercle at base of first stria distinct, with just traceable tubercle at base of third interval, basal declivity with isodiametric reticulation. Humeral tooth small, situated at base of fourth interval. Striae moderately deep, narrow, distinctly and regularly punctuate-striate, one to three free at base, four to six reaching humerus, one and two indistinctly joining at apex, three and four, and five and six distinctly joining apically. Intervals convex, the two lateral ones more convex, interval eight narrowed apically. Surface of intervals glossy, interval eight with isodiametric reticulation in males and females.

Hind wing: In one of the two females, the hind wing is reduced to half of the length and width of the elytron. In the other female and the male the hind wing is not traceable.

Lower surface: Proepisternum in lateral half distinctly isodiametrically reticulated, with few transverse wrinkles. Sternite of abdomen in both sexes smooth and glossy at middle, with moderately sized punctures laterally, in females in addition with traces of isodiametric reticulation baso-laterally. The two marginal setigerous punctures are widely separated.

Legs: Protibia covered with distinct longitudinal reticulation, with the three spines of moderate length, not sulcate dorsally, movable spur nearly as long as apical spine, slightly arcuate in apical third. Intermediate tibia with six tubercles at the inner side and eight at the lateral side, tubercles furnished with strong setae.

Male genitalia ( Fig. 9 ): Median lobe massive, moderately short in comparison to body size, distinctly arcuate at middle, apex straight, with flattened, asymmetric acuminated and slightly contorted, with dorsal bulge in apical half. Endophallus densely folded, with group of bristles. Ventral paramere with apophysis, somewhat distorted, apex unknown. Dorsal paramere sinuslike arcuate, with widened apophysis, apex with four long setae

Female gonocoxite and epipleurite ( Fig. 14 ): Gonocoxite foliform, widened, flattened dorsally and ventrally, with three long nematiform setae positioned at the median side and arising from obtuse tubercles, with one long apical nematiform seta arising from the top of a short protuberance. Epipleurite semi-rectangular, with conspicuously widened angulated rod.

Sexual dimorphism at the integument: Not observed.

VARIATION. In the three available specimens, the triangle-like impressions at the middle of the frons varies in size. On the elytron, stria one and two are joining at apex at one side in two specimens.

DISTRIBUTION. Known from Tanganyika, (continental part of Tanzania).

REMARK. It should be mentioned that E. schmidli sp. nov. is similar to Sulciclivina attenuata ( Herbst, 1806) from the Oriental region. By contrast, it differs by the less swollen proepisternum, the missing punctures on the disk of the pronotum, the presence of an interruption of the umbilical setigerous punctures in the lateral channel of the elytron, the female gonocoxites which are stouter, have only three setae at the median side, a short tubercle at the apex, and an additional seta at the inner side on the surface of the gonocoxites. The male aedeagus is not bisinuate curved at the apex and the apical lamella is more acuminated and hollowed out. The character differences between Eoclivina and Sulciclivina have been keyed out in BALKENOHL (2022).

Figures

Figs 1-5. Eoclivina species, habitus, dorsal view. 1, E. dumolinii (Putzeys, 1846), ♂, specimen from Velingara, Guinea. 2, E. basilewskyi sp. nov., holotype. 3, E. machadoi (Basilewsky, 1955), holotype. 4, E. burgeoni (Kult, 1959), holotype. 5, E. schmidli sp. nov., holotype.

Figs 6-9. Eoclivina species, male genitalia. A-B, aedeagus. A, dorsal view. B, lateral view. C-D, parameres, dorsal view. 6, E. dumolinii (Putzeys, 1846), specimen from Velingara, Guinea. 7, E. basilewskyi sp. nov., holotype. 8, E. machadoi (Basilewsky, 1955), paratype from Riv. Chiumbe, Angola. 9, E. schmidli sp. nov., holotype.

Figs 10-14. Eoclivina species, females, gonocoxite and epipleurite, ventro-lateral view. 10, E. dumolinii (Putzeys, 1846), specimen from Velingara, Guinea. 11, E. basilewskyi sp. nov., paratype. 12, E. machadoi (Basilewsky, 1955), paratype. 13, E. burgeoni (Kult, 1959), holotype. 14, E. schmidli sp. nov., paratype.