Eoclivina machadoi ( BASILEWSKY, 1955 )

Balkenohl, Michael, 2023, The genus Eoclivina (KULT, 1959) from Africa with the description of two new species (Coleoptera, Carabidae, Clivinini), Belgian Journal of Entomology 138, pp. 1-17 : 12-13

Eoclivina machadoi ( BASILEWSKY, 1955)

( Figs 3 , 8 , 12 )

Clivina machadoi BASILEWSKY, 1955: 101 ;

Clivina (Eoclivina) machadoi ( BASILEWSKY, 1955) , KULT, 1959: 215; LORENZ 2005: 145; 2023: 5 ZCCZ;

Eoclivina machadoi ( BASILEWSKY, 1955) , BALKENOHL, 2022: 108.

TYPE MATERIAL. Holotype: ♀, with white black printed and handwritten in black ink label and data: “ Angola: Dundo Forêt (Luachimo) IX. 1948 A.deBarrosMachado” / handwritten in black ink: “11 30.4” / red, black framed and printed: “HOLOTYPUS” / white, handwritten in black ink and printed: “ Clivina machadoi , n.sp. P. Basilewsky det., 1953” / white, black printed: “RMCA ENT 000019856” and a bar code ( Fig. 3 ) ( MRACT).

Paratypes: 1 ♂, same data as holotype ( MRACT) ; 1 ♂, 1 ♀, with white black printed label and data: “ Angola: Riv. Chiumbe, 40 km E. de Dundo V-1949 A.deBarrosMachado” / handwritten in black ink: “1493.18” / red, black framed and printed: “ PARATYPUS ” / white, handwritten in black ink and printed: “ Clivina machadoi , n.sp. P. Basilewsky det., 1954” ( MRACT) .

ADDITIONNAL MATERIAL. ANGOLA. 1 ♂, 1 ♀, Angola : 15908.6 7.20 S, 20.52 E; alt. 720 m R. Cambuacala, env. Dundo A. de Barros Machado ( MRAC) ; 2 ♂♂, ANG. 14614.14 ( MRAC) . DEMOCRATIC REPUBLIC OF THE CONGO. 1 ♀, Coll. I.R.Sc.N.B. Congo D.R. Kona 2°02’32.97”N, 22°47’26.09”E 11.v.2010, swamp forest Light trap, leg. P. Grootaert I.G.: 33.593 ( RBINS) .

EXTENDED DIAGNOSIS. A medium sized piceous species with the following measurements: Body length 5.1–5.53 mm (x = 5.39 mm *), width 1.51–1.61 mm (x = 1.58 mm *), ratio L/W of pronotum 0.95–0.99 (x = 0.97*), ratio L/W of elytra 1.82–1.87 (x = 1.84*); (*n = 9). The elytron is unicolor. Wing of the clypeus relatively wide but nearly fused with lateral lobe of the clypeus. Eye convex and projecting laterally. Pronotum with lateral margin slightly attenuating anteriorly with straight parts anterior middle, on the disk with scattered small to moderate sized punctures, punctures larger towards base, proepisternum just not visible in dorsal view. Elytra with the shape long-oval with straight and diverging part of the lateral margins, with maximum width slightly behind middle, intervals with different pattern of reticulation among the sexes: In males, interval eight reticulated, all the others are smooth and glossy. In females, the reticulation covers intervals one, and six to eight. In addition, the base is reticulated as well as around a fifth of the apex. Intervals two to five smooth on the disk. Row of umbilical setigerous punctures of the lateral channel interrupted posterior humerus and at middle by one puncture. Abdominal sternite in males with minute punctures, smooth and glossy at middle, laterally with some smaller punctures, sternite of females distinctly and completely covered with transverse reticulation. Intermediate tibia with six tubercles at the inner side and eight at the lateral side, tubercles furnished with strong setae.

Hind wing: Fully developed. Male genitalia ( Fig. 8 ): Median lobe wide in lateral view, slightly arcuate, with small and rounded spatula. Endophallus with longitudinal group of bristles apically.

Female gonocoxite and epipleurite ( Fig. 12 ): Gonocoxite foliform, widened, with four robust nematiform setae positioned at the median side, apically with one long seta. Epipleurite quadrangular, asetose, with widened angulated rod.

Sexual dimorphism at the integument: Males and females differ considerably in the reticulation on the intervals of the elytron and the abdominal sternites (see diagnosis above).

VARIATION. On the disk of the pronotum, the punctures are more or less dense and vary in its distinctness. In some of the females, the reticulation on the intervals of the elytron is not strongly expressed on interval six. In others, it extends slightly to interval two and five.

The distinguishing character from the similar species E. burgeoni KULT is the reticulation on the intervals of the elytron. In the female holotype of E. burgeoni , all intervals are completely reticulated.

DISTRIBUTION. The species is found in the North and West of Angola, and in the Democratic Republic of the Congo.

Figures

Figs 1-5. Eoclivina species, habitus, dorsal view. 1, E. dumolinii (Putzeys, 1846), ♂, specimen from Velingara, Guinea. 2, E. basilewskyi sp. nov., holotype. 3, E. machadoi (Basilewsky, 1955), holotype. 4, E. burgeoni (Kult, 1959), holotype. 5, E. schmidli sp. nov., holotype.

Figs 6-9. Eoclivina species, male genitalia. A-B, aedeagus. A, dorsal view. B, lateral view. C-D, parameres, dorsal view. 6, E. dumolinii (Putzeys, 1846), specimen from Velingara, Guinea. 7, E. basilewskyi sp. nov., holotype. 8, E. machadoi (Basilewsky, 1955), paratype from Riv. Chiumbe, Angola. 9, E. schmidli sp. nov., holotype.

Figs 10-14. Eoclivina species, females, gonocoxite and epipleurite, ventro-lateral view. 10, E. dumolinii (Putzeys, 1846), specimen from Velingara, Guinea. 11, E. basilewskyi sp. nov., paratype. 12, E. machadoi (Basilewsky, 1955), paratype. 13, E. burgeoni (Kult, 1959), holotype. 14, E. schmidli sp. nov., paratype.