Entocythere (Cytherites) riojai, Williams & Weaver, 2018

Williams, Bronwyn W. & Weaver, Patricia G., 2018, A historical review of the taxonomy and classification of Entocytheridae (Crustacea: Ostracoda: Podocopida), Zootaxa 4448 (1), pp. 1-129 : 83

* Thermastrocythere riojai ( Hoff, 1943) Hobbs & Hobbs, 1970

1943 Entocythere (Cytherites) riojai , n. sp. —Hoff: 276–282, 286, Fig. 1A–L (D, E, H, R, T).

1943a Entocythere (Cytherites) riojai Clayton Hoff. — Rioja: 564, 565 (R).

1943b Entocythere (Cytherites) riojai Clayton Hoff (1943) .— Rioja: 567, 582, 583 (R, T).

1944a E. riojai Hoff 1943 .—Hoff: 332, 337 (K, R).

1945 Entocythere riojai Clayton Hoff. — Rioja: 422 (R).

1947 E. riojai Hoff. —Tressler: 705 (C).

1949 Entocythere riojai Hoff. — Rioja: 328 (R).

1954 E. riojai Hoff (1943) .—Kozloff & Whitman: 162 (R).

1955 E. roijai Hoff (1943) .—Hobbs: 330, erroneous spelling (R).

1959 E. riojai Hoff (1943) .—Crawford: 167 (C).

1959 Entocythere riojai Hoff 1943 .—Tressler: 730, Fig. 28.188a, b, c, d (D, H, K, T).

1961 E. riojai Hoff (1943) .—Crawford: 238 (R).

1962 Uncinocythere riojai (Hoff) new combination.—Hart: 122 [as E. riojai ], 138 (C, D, T).

1966 Thermastrocythere harti new species.—Hobbs & Walton: 8–9, 10, 11, Fig. 2e–g (D, H, R, T).

1969a Thermastrocythere harti Hobbs and Walton, 1966 . — Hart & Hart: 185 (D, R, T).

1970 Thermastrocythere harti Hobbs, Jr. & Walton, 1966 .—Delorme: 1257, 1258 [by implication] (C, D, H).

1970 Thermastrocythere riojai ( Hoff, 1943) .—Hobbs & Hobbs: 17 [and as Entocythere riojai and Thermastrocythere harti Hobbs and Walton, 1966:8 ], new combination (R, T).

1971 Thermastrocythere rioja ( Hoff, 1943) .—Hobbs III: 143, erroneous spelling (A).

1974 Thermastrocythere riojai ( Hoff, 1943) .—Hart & Hart: 14 [as E. riojai ], 117 [and as Entocythere riojai , Thermastrocythere harti , and Entocythere harti , the latter lapsus calami], 118–120 [and as Entocythere riojai and Thermastrocythere harti ], 210, 230, Pl. XXXIV Figs. 3–5, Pl. LIV (C, D, H, R, T).

1975 Thermastrocythere riojai ( Hoff, 1943) .—Hart & Hart: 44–45 [and as Thermastrocythere harti (= Entocythere riojai )] (C, D, H).

1990 Thermastrocythere riojai (Hoff) .—Clamp: 14 (D, E, H).

2011 Thermastrocythere riojai ( Hoff, 1943) .—Williams et al.: 276–279, 281–282, Fig. 1 (C, D, H).

2014a Thermastrocythere riojai ( Hoff, 1943) .—Mestre, Monrós & Mesquita-Joanes: 950 (C).

Remarks: Hoff (1943: 276) designated a biunguis female in copulation as the holotype and the adult male of the pair as the allotype, indicating that several of the paratypes are females, both gravid (= triunguis) and copulating (= biunguis). Further, the majority of the description is dedicated to the female. This type designation was repeated for all species described in Hoff (1942b) and Hoff (1943); Hoff (1944a) shifted to what would become convention, designating a male as the holotype and a female as the allotype. The subgenus designation was removed from Entocythere (Cytherites) riojai in Hoff (1944a) . Thermastrocythere harti was designated by Hobbs & Walton (1966) as the type species for the newly erected genus Thermastrocythere . While examining type material deposited by C. Clayton Hoff in the USNM Collection, Horton H. Hobbs, Jr. and Dabney G. Hart found that the paratypes of Entocythere riojai were in fact two species. The holotype (= allotype by designation) and a partially dissected paratypic (?) male of E. riojai were found to be nearly identical to the holotype of T. harti , making this latter species a subjective junior synonym to Hoff’s E. riojai . The synonymization was reported in Hobbs & Hobbs (1970: 16–17).

Figures

FIGURE 1. Ankylocythere ancyla (NCSM 29785), adult male, from Bladen County, North Carolina, USA. Inset shows a magnified view of a terminal claw of a thoracic leg, specifically, a fifth appendage.