Ensiosmylus Khramov, 2014a

Winterton, Shaun L., Martins, Caleb Califre, Makarkin, Vladimir, Ardila-Camacho, Adrian & Wang, Yongjie, 2019, Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera, Zootaxa 4581 (1), pp. 1-99 : 60

Ensiosmylus Khramov, 2014a

Type species. Ensiosmylus acutus Khramov, 2014a: 302 (by original designation).

Diagnosis. Hind wing narrow, slightly falcate; subcostal veinlets relatively widely spaced; stem of HW RP relatively long, RP1 originates far from origin of RP; RP with 17 branches, slightly sinuous distally, crossveins semi-regular in arrangement with distinct outer gradate series present; HW MA with very few dichotomous branches close to wing margin, MP with five pectinate branches; HW CuA with ten regular pectinate branches, with few forks, CuP with single terminal fork; A1 with three pectinate branches; FW unknown.

Comments. Khramov (2014a) assigned this genus to Spilosmylinae based on the branching pattern of the anal veins and crossvein pattern in the radial and medial areas. However, a typical feature of Spilosmylinae (i.e., the presence of a spur-like process on the basal part of the hind wing MP directed proximally) is not discernible in the fossil. The hind wing figures presented by Khramov (2014a) have the veins identified following Kukalov-Peck & Lawrence (2004) (i.e., MA labelled as MP, MP as CuA, CuA as CuP and so on). The presence of a simple CuP vein in hind wing could place Ensiosmylus in Spilosmylinae , but the feature also occurs in Protosmylinae and Gumillinae ; consequently, its placement in Spilosmylinae is tentative until a more complete specimen is known; placement in Gumillinae may be precluded by the presence of a well-define outer gradate series.