Elmomorphus hamatus, Selnekovič & Jäch & Kodada, 2024

Selnekovič, Dávid, Jäch, Manfred A. & Kodada, Ján, 2024, Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae), European Journal of Taxonomy 957, pp. 1-229 : 195-199

Elmomorphus hamatus sp. nov.

urn:lsid:zoobank.org:act:72E22E5E-4709-4E97-BF2B-32CF9CC6467F

Figs 96–97 , 106A , 115C

Differential diagnosis

Elmomorphus hamatus sp. nov. ( Fig. 96 ) is characterised by the strongly convex body and by having the dorsal plastron confined to small areas on the frontoclypeus around the antennal insertions. The elytra has small, scattered punctures and shallow, large punctures arranged in nine shallow striae. The head is microreticulate in anterior portions, the pronotum is smooth, and the elytra weakly microreticulate. Elmomorphus hamatus most closely resembles E. superficialis sp. nov., but differs in its larger body dimensions (TL in E. hamatus : ♂♂ 3.58–4.11 mm (3.75 ±0.14, n= 10), ♀♀ 3.74–4.00 mm (3.88 ± 0.08, n= 10), in E. superficialis : ♂♂ 3.21–3.52 mm (3.36 ±0.10, n=13), ♀♀ 3.24–3.75 mm (3.49 ± 0.14, n= 13)), less convex and more elongate body, punctures on the pronotal disc being distinctly smaller than those on the head, and the parameres being strongly curved apically ( Fig. 97 ). From E. simplex sp. nov., E. glabriclunis sp. nov., and E. comosiclunis sp. nov. it differs in the presence of superficial elytral striae. From E. jii sp. nov., E. schoenmanni sp. nov., and E. vietnamensis sp. nov. it can be distinguished by the presence of plastron on the entire first abdominal ventrite and by the absence of plastron in the posteromedian portion along the posterior margin of ventrite 5.

Etymology

The epithet ‘hamatus’ (‘hooked’) is a Latin adjective in the nominative singular and refers to the curved parameres.

Type material

Holotype

CHINA – Hunan Province • ♂; “ CHINA: Hunan, 30.3.2003 Dawei Shan Nat. Park 114°06′31″E 28°25′01″N ca. 1450m, leg. Schönmann, Komarek & Wang (CWBS 519)”; IAECAS.

Paratypes

CHINA – Hunan Province • 3 ♂♂, 6 ♀♀, 3 ex.; same collection data as for holotype; CKB, NMW • 7 ♂♂, 7 ♀♀, 1 ex.; “ CHINA: Hunan, 30.3.2003 Dawei Shan (outside NP) 114°11′10″E 28°25′44″N ca. 1360 m leg. Schönmann, Komarek & Wang (CWBS 518)”; CKB, NMW • 4 ex.; “ CHINA: Hunan, 30.3.2003 Dawei Shan Nat. Park 114°06′31″ E 28°25′01″ N ca. 1450 m, leg. Schönmann, Komarek & Wang (CWBS 520)”; NMW .

Type locality

China, Hunan Province, Dawei Mountain National Forest Park, 28°25′1″ N, 114°6′31″ E.

Description

Measurements (mm): TL: ♂♂ 3.58–4.11 (3.75± 0.14, n= 10), ♀♀ 3.74–4.00 (3.88 ± 0.08, n=10); PL: ♂♂ 0.94–1.04 (0.95± 0.03, n =10), ♀♀ 0.87–0.98 (0.94 ± 0.03, n=10); PW: ♂♂ 1.73–1.98 (1.83± 0.07, n =10), ♀♀ 1.77–1.94 (1.86 ±0.05, n =10); EL: ♂♂ 2.71–3.13 (2.84± 0.12, n =10), ♀♀ 2.92–3.13 (3.00 ±0.08, n=10); EW: ♂♂ 1.92–2.21 (2.04 ± 0.09, n= 10), ♀♀ 2.00–2.17 (2.08 ± 0.05, n= 10); PhL: 0.55–0.61 (0.58 ± 0.02, n=10); PrL: 0.39–0.44 (0.41 ± 0.01, n=10).

Body oval, strongly convex, widest just before elytral midlength ( Fig. 96 ). Colouration black, antennae, mouthparts, and legs reddish brown. Dorsal pubescence consists of short, thin, decumbent setae arising from small, round punctures. Dorsal plastron present only on small areas of frontoclypeus around antennal insertions. Ventral plastron on entire ventral surface, except for prosternal process, metaventrite in median portion, and abdominal ventrite 5 in posteromedian portion and along posterior margin.

Cranium with small, round punctures, each smaller than eye facet, separated from each other by ca 0.5–1.0 × puncture diameter; plastron present in small lateral areas of frontoclypeus around antennal insertions. Labrum transverse, anterior margin straight, exposed portion microreticulate, with small setiferous punctures; setae equal in length in both sexes. Anterior margin of clypeus straight. Eyes oval, weakly protruding, ID: ♂♂ 0.61–0.72 mm (0.64 ± 0.03, n =10), ♀♀ 0.66–0.70 mm (0.67 ± 0.02, n =10). Antennae short, 11-segmented, densely setose.

Pronotum transverse, moderately convex, widest at base, PW/PL: ♂♂ 1.84–2.00 (1.92± 0.06, n=10), ♀♀ 1.89–2.05 (1.97 ±0.06, n=10); pronotal disc smooth, with round setiferous punctures smaller than those on head; anterior angles protruding and deflexed; lateral pronotal sides convergent and rounded. Prosternal process with lateral and posterior edges rounded, lateral margins slightly raised, without clusters of erect setae, median keel arched. Scutellum slightly longer than wide, smooth with several setiferous punctures. Metaventral process with lateral margins slightly raised, metaventral disc flattened, area without plastron widened posteriorly. Elytra oval, widest before middle, EL/EW: ♂♂ 1.36–1.45 (1.40 ± 0.02, n= 10), ♀♀ 1.42–1.50 (1.44± 0.03, n=10); plastron absent; surface with very weak microreticulation formed by irregular meshes; small punctures scattered over entire surface, shallow, large punctures hardly discernible, placed in nine indistinct striae. Tibiae slightly curved; protibia ca 1.4× as long as protarsus; PrTL/PL: ♂♂ 0.90–0.98 (0.94± 0.03, n=10), ♀♀ 0.86–1.01 (0.95 ± 0.05, n= 10). Terminal protarsomere ca as long as three preceding segments combined.

Ventrites covered with plastron, except for median area on posterior half of ventrite 5, which expands along posterior margin of ventrite. Ventrite 5 evenly convex in both sexes; posterior margin weakly emarginate in male, rounded in female. Aedeagus ( Fig. 97 ): phallobase short, broad, PhL/PrL: 1.34–1.47 (1.42 ± 0.04, n= 10); parameres widest basally, gradually evenly tapering toward apices, nearly straight along basal two-thirds then strongly curved ventrad at apical third, apices widely rounded (lateral aspect); penis apex narrowly rounded, reaching only slightly over parameral midlength; sclerotised fibula rather long, slender. Bursa copulatrix with two lateral clusters of numerous minute microsclerites in distal part, and with numerous minute ventrolateral microsclerites in proximal part ( Fig. 106A ).

Secondary sexual dimorphism

Ventrite 5 in males weakly emarginated at apex, in females with rounded apical margin.

Distribution

China (Hunan) ( Fig. 115C ).

Figures

Fig. 96. Elmomorphus hamatus sp. nov., holotype, male (IAECAS), TL: 3.75 mm.

Fig. 97. Elmomorphus hamatus sp. nov., holotype (IAECAS), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.

Fig. 106. Bursa copulatrix and vagina. A. Elmomorphus hamatus sp. nov., paratype from type locality (NMW). B. E. oblongus sp. nov., paratype from type locality (NMW). C. E. glabriclunis sp. nov., paratype from China, locality CWBS 258 (CKB). D. E. simplex sp. nov., paratype from type locality (CKB). E. E. comosiclunis sp. nov., paratype from China, locality CWBS 468. Scale bar: 0.1 mm.

Fig. 115. Distribution maps.A. Elmomorphus reticulatus sp. nov.B. E. superficialis sp. nov.C. E. hamatus sp. nov. D. E. oblongus sp. nov. E. E. glabriclunis sp. nov. F. E. simplex sp. nov.