Tesnus stillatus, L. Benick, 1931

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium IV. Staphylinine Group (Part 1) Euaesthetinae, Leptotyphlinae, Megalopsidiinae, Oxyporinae, Pseudopsinae, Solieriinae, Steninae, Bulletin of the American Museum of Natural History 2440 (265), pp. 1807-2440 : 1807-2440

publication ID

https://doi.org/ 10.1206/0003-0090.265.1.4

persistent identifier

https://treatment.plazi.org/id/0F6987E5-FDC1-A018-FF7C-F7A9FCD6F911

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scientific name

Tesnus stillatus
status

 

stillatus L. Benick, 1931 , see: kempi Bernhauer, 1926 .

stomachosus Scheerpeltz, 1933, see: abdominalis Fauvel, 1895 .

strabo Puthz, 1986 j: 301 ( Stenus ; subgenus Hypostenus ; Type locality: Cameroun:

Ft. de Bafut, Nguemba).

- DISTRIBUTION: Cameroon.

strandi L. Benick, 1937b: 183 ( Stenus ; subgenus Nestus ; Type locality: Norway. Also cited from numerous northern European localities).

— Palm, 1961a: 86 ( Stenus ; subgenus Nestus ; characters; Sweden; Norway; Finland).

— Puthz, 1970n: 302 ( Stenus ; note; aedeagus).

— Haberman, 1983: 101 ( Stenus ; Estonia).

— Ryabukhin, 1999: 47 ( Stenus ; subgenus Nestus ; catalog; habitat; distribution; northeastern Russia).

- DISTRIBUTION: Europe, Russia.

strangulatus Casey, 1884: 39 ( Stenus ; Type locality: New York; Detroit , Michigan).

— Blatchley, 1910: 406 ( Stenus ; characters; Indiana).

— Bernhauer and Schubert, 1911: 167 ( Stenus ; catalog).

— Campbell and Davies, 1991: 112 ( Stenus ; Canada).

— Downie and Arnett, 1996: 490 ( Stenus ; characters; USA).

- DISTRIBUTION: USA, Canada.

strejceki Hromádka, 1985: 189 ( Stenus ; subgenus Parastenus ; Type locality: USSR: Abchazia, Caucasus occ. Amtkel pr. Suchumi, 400 m).

— Ryvkin, 1990: 182 ( Stenus ; cited as streiceki; notes; Georgia).

- DISTRIBUTION: Georgia.

striatellus L. Benick, 1932 , see: Dianous .

strigicollis L. Benick, 1949b: 564 ( Stenus ; subgenus Tesnus ; Type locality: Brasilien: Barra da Boa Vista, atual Coripós, Nebenarm des Rio São Francisco, Mun. Boa Vista atual Coripós, Staat Pernambuco; Lagoa dos Canudos, nw. Petrolina, Mun. Petrolina ,

Staat Pernambuco).

- DISTRIBUTION: Brazil.

strigosus Fauvel, 1873: 81 [= 1873a: 239] ( Stenus ; Type locality: Corse, L’Ospidale ; [Note: See lectotype designation by Puthz, 1969]).

— Sainte-Claire Deville, 1906: 80 ( Stenus ; Corsica).

— Bernhauer and Schubert, 1911: 167 ( Stenus ; catalog).

— L. Benick, 1929a: 33 ( Stenus ; subgenus Nestus ; characters in key).

— Portevin, 1929: 398 ( Stenus ; characters; France).

— Scheerpeltz, 1933: 1166 ( Stenus ; subgenus Stenus ; catalog).

— Puthz, 1967f: 272 ( Stenus ; characters; Morocco).

— Puthz, 1969: 14 ( Stenus ; lectotype designation; notes).

— Puthz, 1970l: 206 ( Stenus ; characters; Sardinia).

— Dauphin, 1993: 178, 183 ( Stenus ; subgenus Nestus ; characters; Corsica; Sardinia).

— Puthz and Zanetti, 1995: 19 ( Stenus ; Italy).

— Marchal and Chardonnet, 2000: 71 ( Stenus ; Corsica).

- DISTRIBUTION: Morocco, France, Italy.

strupiianus Scheerpeltz, 1958, see: hypsidromus Ganglbauer, 1902 .

stygicus Say, 1831: 45 [= 1834: 458] ( Stenus ; Type locality: Missouri).

— Erichson, 1840: 698 ( Stenus ; characters; America septentrionalis).

— LeConte, 1850: 220 ( Stenus ; Lake Superior).

Casey, 1884: 104 ( Stenus ; characters; Lake Superior; Texas; Kansas; western States; Alabama; southern States ; middle States; Michigan; New Jersey; District of Columbia; Pennsylvania; New York; Massachusetts) .

— Blatchley, 1910: 409 ( Stenus ; characters; Indiana).

— Bernhauer and Schubert, 1911: 167 ( Stenus ; catalog).

— Sanderson, 1957: 262 ( Stenus ; subgenus Stenus ; characters; Washington).

— Puthz, 1972b: 12 ( Stenus ; characters; Michigan).

— Puthz, 1988b: 141 ( Stenus ; characters; California; Utah; Nevada; Mississippi; Kentucky).

— Campbell and Davies, 1991: 112 ( Stenus ; USA; Canada).

— Dearborn and Donahue, 1994: 14 ( Stenus ; Maine).

— Downie and Arnett, 1996: 491 ( Stenus ; characters; USA).

- DISTRIBUTION: USA, Canada.

suaveolus Sharp, 1886: 667 ( Stenus ; Type locality: Guatemala, Paso Antonio ).

— Bernhauer and Schubert, 1911: 178 ( Stenus ; subgenus Hypostenus ; catalog).

— L. Benick, 1938c: 277 ( Stenus ; characters).

- DISTRIBUTION: Guatemala.

subaeneus Erichson, 1840: 727 ( Stenus ; Type locality: Lutetiae; [Note: See lectotype designation by Puthz, 1967h]. Also cited from Gallia meridionalis; Sardinia; Anglia ).

— Leprieur, 1851: 199 ( Stenus ; characters).

— Kiesenwetter, 1851a: 426 ( Stenus ; Spain).

— Waterhouse and Janson, 1855: 147 ( Stenus ; synonym of gonymelas ).

— Fairmaire and Laboulbène, 1856: 592 ( Stenus ; characters; France).

— Redtenbacher, 1857: 989 ( Stenus ; characters; Austria).

— Kraatz, 1857d: 786 ( Stenus ; characters; Germany).

— Kraatz, 1858b: 125 ( Stenus ; Crete; Greece).

— G. Waterhouse, 1858: 29 ( Stenus ; synonym of gonymelas ).

— Rye, 1864: 91 ( Stenus ; synonym of gonymelas ).

— Fauvel, 1873: 123 [= 1873a: 281] ( Stenus ; characters; France; Britain; Germany; Switzerland; Italy; Corsica; Sardinia; Crete; Algeria).

— Redtenbacher, 1874: 246 ( Stenus ; characters).

— Fauvel, 1878: 103 [= 1878a: 23] ( Stenus ; Algeria).

— Rey, 1884: 334 [= 1884a: 182] ( Stenus ; subgenus Mesostenus ; characters; France).

— Fauvel, 1886: 31 [= 1886a: 23] ( Stenus ; Algeria).

— Fowler, 1888: 347 ( Stenus ; characters; Britain).

— Ganglbauer, 1895: 594 ( Stenus ; subgenus Mesostenus ; characters; western middle Europe; Mediterranean region).

— Fauvel, 1902b: 78 ( Stenus ; Algeria).

— Sainte-Claire Deville, 1906: 83 ( Stenus ; Corsica).

— Reitter, 1909: 160 ( Stenus ; subgenus Parastenus ; characters; Germany).

— Bernhauer and Schubert, 1911: 185 ( Stenus ; subgenus Hemistenus ; catalog).

— Fiori, 1915: 8 ( Stenus ; Italy).

— L. Benick, 1929a: 68 ( Stenus ; subgenus Parastenus ; characters in key).

— Portevin, 1929: 403 ( Stenus ; characters; France).

— Scheerpeltz, 1933: 1197 ( Stenus ; subgenus Hemistenus ; catalog).

— Wüsthoff, 1934a: Tafel III ( Stenus ; illustration of aedeagus).

— Normand, 1935: 364 ( Stenus ; Algeria; Tunisia).

— Tottenham, 1949: 413 ( Stenus ; checklist; Britain).

— Jeannel and Jarrige, 1949: 348 ( Stenus ; cave species; Spain).

— Scheerpeltz, 1961d: 122 ( Stenus ; subgenus Parastenus ; Italy).

— Szujecki, 1961: 61 ( Stenus ; subgenus Parastenus ; characters; Poland).

— Horion, 1963: 388 ( Stenus ; subgenus Parastenus ; Britain; Holland; Belgium; France; Switzerland; Spain; Italy; Dalmatia; Greece; Bulgaria; Hercegovina; Macedonia; Turkey; Austria; Germany).

— Lohse, 1964: 125 ( Stenus ; subgenus Parastenus ; characters; central Europe).

— Focarile, 1964: 63 ( Stenus ; Italy).

— Scheerpeltz, 1965b: 396 ( Stenus ; Sicily).

— Scheerpeltz, 1965c: 501 ( Stenus ; Sicily).

— Puthz, 1966f: 122 ( Stenus ; subgenus Parastenus ; notes).

— Puthz, 1967: 9 ( Stenus ; subgenus Parastenus ; Portugal).

— Puthz, 1970p: 177 ( Stenus ; Italy).

Puthz, 1971v: 87 ( Stenus ; note; Turkey; Lebanon) .

— Puthz, 1972s: 274 ( Stenus ; Yugoslavia; Albania; Greece).

— Tikhomirova, 1973a: 174 ( Stenus ; subgenus Parastenus ; checklist; USSR).

— Pope, 1977: 26 ( Stenus ; Britain).

— Puthz, 1980e: 366 ( Stenus ; Albania).

— Waterston, Owen, Welch, Bacchus and Hammond, 1981: 274 ( Stenus ; Outer Hebrides).

— Puthz, 1983e: 350 ( Stenus ; notes; Georgia).

— Seger, 1986: 34 ( Stenus ; Belgium).

— Lucht, 1987: 98 ( Stenus ; checklist; Germany; Austria; Switzerland; France; Benelux).

— Gusarov, 1989: 7 ( Stenus ; habitat notes; Ukraine).

— Ryvkin, 1990: 207 ( Stenus ; notes; Russia; Georgia).

— Welch, 1993: 228 ( Stenus ; ovariole number and ovary structure).

— Puthz and Zanetti, 1995: 19 ( Stenus ; Italy).

— Herman, 2001: 34 ( Stenus ; nomen protectum).

- DISTRIBUTION: Algeria, Europe, Russia, Georgia, Turkey, Lebanon.

geniculatus Mannerheim, 1830: 43 ( Stenus ; [preoccupied]; Type locality: E. Gallia meridionalis).

— Mannerheim, 1831: 457 ( Stenus ; characters; France).

— Erichson, 1840: 727 ( Stenus ; synonym of subaeneus ).

— Fairmaire and Laboulbène, 1856: 592 ( Stenus ; synonym of subaeneus ).

— Kraatz, 1857d: 786 ( Stenus ; synonym of subaeneus ).

— Rey, 1884: 335 [= 1884a: 183] ( Stenus ; synonym of subaeneus ).

— Ganglbauer, 1895: 594 ( Stenus ; synonym of subaeneus ).

— Bernhauer and Schubert, 1911: 185 ( Stenus ; synonym of subaeneus ).

gonymelas Stephens, 1833: 291 ( Stenus ; [Note: This name is older than subaeneus ]; Type locality: London; Bristol; Norfolk; Suffolk; Devonshire; Crwmlyn Bog; Cramond).

— Stephens, 1839: 413 ( Stenus ; characters; England).

— Waterhouse and Janson, 1855: 147 ( Stenus ; synonyms).

— Rye, 1864: 91 ( Stenus ; characters; England).

— Fauvel, 1873: 123 [= 1873a: 281] ( Stenus ; synonym of subaeneus ).

— Fowler, 1888: 347 ( Stenus ; synonym of subaeneus ).

— Ganglbauer, 1895: 594 ( Stenus ; synonym of subaeneus ).

— Bernhauer and Schubert, 1911: 185 ( Stenus ; synonym of subaeneus ).

— Herman, 2001: 34 ( Stenus ; nomen oblitum).

geniculatus Stephens, 1833: 291 ( Stenus ; [preoccupied]; Type locality: London; Devonshire).

— Stephens, 1839: 412 ( Stenus ; characters; England).

— Waterhouse and Janson, 1855: 147 ( Stenus ; synonym of gonymelas ).

— G. Waterhouse, 1858: 29 ( Stenus ; synonym of gonymelas ).

subarcticus Poppius, 1908a: 106 ( Stenus ; subgenus Nestus ; Type locality: Lutto-Fluss in der Nähe von Komsiovaara).

— Bernhauer and Schubert, 1911: 167 ( Stenus ; catalog).

— Puthz, 1966e: 301 ( Stenus ; subgenus Nestus ; subspecies of gibbicollis ; characters).

— Puthz, 1972e: 107 ( Stenus ; valid species).

- DISTRIBUTION: Norway, Finland.

bernhaueri Poppius, 1908a: 107 ( Stenus ; subgenus Nestus ; Type locality: Lutto-Fluss in der Nähe von Komsiovaara).

— Bernhauer and Schubert, 1911: 154 ( Stenus ; catalog).

— L. Benick, 1921c: 193 ( Stenus ; synonym of gibbicollis ).

— Puthz, 1972e: 107 ( Stenus ; synonym of subarcticus ).

polaris Münster, 1911: 280 ( Stenus ; Type locality: Flusse Gorzzejok in Karasjok im nördlichen Norwegen).

— L. Benick, 1921c: 193 ( Stenus ; synonym of gibbicollis ).

subascendens Puthz, 1967 i: 222 ( Stenus ; subgenus Hypostenus ; Type locality: Dar-Banda mérid., Krébédjé, Fort Sibut).

— Puthz, 1971: 300 ( Stenus ; subgenus Hypostenus ; characters; Central African Republic).

- DISTRIBUTION: Chad, Central African Republic.

subcautus Ryvkin, 2000: 350 ( Stenus ; subgenus Nestus ; Type locality: Martime Province, Khasan, Golubinyi Utyos).

- DISTRIBUTION: Russia.

subconcentus L. Benick, 1921, see: infuscatus L. Benick, 1916 .

subconvexus Rey, 1884, see: salinus Brisout, 1863 .

subcoeruleus Bernhauer, 1916f: 176 ( Stenus ; subgenus Hypostenus ; Type locality: Westliches Kolumbien: Umgebung von Cali am Cauca, Caldas, 1000 m).

— L. Benick, 1928d: 43 ( Stenus ; characters).

— Scheerpeltz, 1933: 1183 ( Stenus ; subgenus Hypostenus ; cited as subcaeruleus; catalog).

— L. Benick, 1938c: 279 ( Stenus ; characters).

— Puthz, 1974a: 68 ( Stenus ; characters; Venezuela).

- DISTRIBUTION: Colombia, Venezuela.

subcordatus Puthz, 1972 s: 277 ( Stenus ; subgenus Parastenus ; Type locality: Dudhatoli,

9000 feet, Gashwal, U. P.; [Note: Gashwal, misspelling for Garhwal]).

- DISTRIBUTION: India.

subcyanescens Bernhauer, 1926a: 126 ( Stenus ; subgenus Hypostenus ; Type locality: Mindanao: Surigao).

— L. Benick, 1929: 245 ( Stenus ; subgenus Hypostenus ; characters; Philippines).

— Scheerpeltz, 1933: 1183 ( Stenus ; subgenus Hypostenus ; catalog).

- DISTRIBUTION: Philippines.

subcylindricus Scriba, 1870: 83 ( Stenus ; Type locality: Santas Albas).

— Fauvel, 1873: 130 [= 1873a: 288] ( Stenus ; characters).

— Bernhauer and Schubert, 1911: 185 ( Stenus ; subgenus Hemistenus ; catalog).

— L. Benick, 1917c: 61 ( Stenus ; characters; notes).

— L. Benick, 1929a: 60 ( Stenus ; subgenus Parastenus ; characters in key).

— Scheerpeltz, 1933: 1197 ( Stenus ; subgenus Hemistenus ; catalog).

— Puthz, 1967: 10 ( Stenus ; subgenus Parastenus ; not in Portugal).

— Puthz, 1967g: 535 ( Stenus ; characters; Spain).

- DISTRIBUTION: Spain.

subdepressus Mulsant and Rey, 1861: 158 [= 1862: 142] ( Stenus ; Type locality: Avenas, montagnes du Beaujolais).

— Fauvel, 1873: 98 [= 1873a: 256] ( Stenus ; characters; France).

— Rey, 1884: 276 [= 1884a: 124] ( Stenus ; subgenus Nestus ; characters; France).

— Ganglbauer, 1895: 575 ( Stenus ; subgenus Nestus ; characters; France; Germany; Austria; Hungary; Spain).

— Reitter, 1909: 158 ( Stenus ; subgenus Nestus ; characters; Germany).

— Bernhauer and Schubert, 1911: 167 ( Stenus ; catalog).

— Bondroit, 1913a: 378 ( Stenus ; synonym of foveiventris ).

— L. Benick, 1917c: 56 ( Stenus ; notes; Germany).

— L. Benick, 1923a: 220 ( Stenus ; notes).

— L. Benick, 1929a: 52 ( Stenus ; subgenus Nestus ; characters in key).

— Portevin, 1929: 399 ( Stenus ; characters; France).

— Linke, 1929: 65 ( Stenus ; ecological notes; Germany; Austria; Romania; Italy; France; Spain; Russia; Poland; Czechoslovakia).

— Donisthorpe, 1930: 99 ( Stenus ; characters; Britain).

— Scheerpeltz, 1933: 1166 ( Stenus ; subgenus Stenus ; catalog).

— Wüsthoff, 1934a: Tafel II ( Stenus ; illustration of aedeagus).

— Szujecki, 1961: 39 ( Stenus ; subgenus Nestus ; characters; Poland).

— Lohse, 1964: 120 ( Stenus ; subgenus Nestus ; characters; central Europe).

— Puthz, 1966f: 118 ( Stenus ; subgenus Nestus ; Germany).

— Puthz, 1971v: 96 ( Stenus ; notes).

— Puthz, 1972s: 257 ( Stenus ; Hungary; Romania).

— Pope, 1977: 26 ( Stenus ; Britain).

— Lucht, 1987: 96 ( Stenus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux).

— Dauphin, 1993: 182, 190 ( Stenus ; subgenus Nestus ; characters; France).

— Hromádka, 1993: 44 ( Stenus ; subgenus Nestus ; Czech Republic).

— Schillhammer, 1994a: 292 ( Stenus ; Austria).

— Puthz and Zanetti, 1995: 19 ( Stenus ; Italy).

- DISTRIBUTION: Europe.

explorator Fauvel, 1873: 96 [= 1873a: 254] ( Stenus ; Type locality: Saint Germain; [Note: See lectotype designation by Puthz, 1969]. Originally cited from Paris, Saint Germain ; Calvados; Plainville; Finistère; Limoges; Biscarosse; Hautes Pyrénées,

vallée de Barousse; Espagne).

— Rey, 1884: 276 [= 1884a: 124] ( Stenus ; synonym of subdepressus ).

— Ganglbauer, 1895: 575 ( Stenus ; synonym of subdepressus ).

— Bernhauer and Schubert, 1911: 167 ( Stenus ; synonym of subdepressus ).

— Puthz, 1969: 14 ( Stenus ; synonym of subdepressus ; lectotype designation: St. Germain).

subditus L. Benick, 1920: 13 ( Stenus ; Type locality: Kaukasus: Daghestan; Martkopi; Helenendorf ).

— L. Benick, 1929a: 72 ( Stenus ; subgenus Parastenus ; characters in key).

— Scheerpeltz, 1933: 1197 ( Stenus ; subgenus Hemistenus ; catalog).

— Puthz, 1967c: 113 ( Stenus ; characters; Caucasus; Daghestan).

— Puthz, 1983e: 349 ( Stenus ; notes; Azerbaidzan; Georgia; Daghestan; Armenia).

— Ryvkin, 1990: 198 ( Stenus ; notes; Russia; Georgia; Azerbaijan).

- DISTRIBUTION: Russia, Georgia, Armenia, Azerbaijan.

subegenus Puthz, 1988b: 151 ( Stenus ; subgenus Nestus ; Type locality: California: Humboldt Co.: Fort Seward).

- DISTRIBUTION: USA.

subescensus Puthz, 1976: 2 ( Stenus ; subgenus Nestus ; Type locality: Nepal: Westnepal, Gebiet des Rarasees, Mahidoela-Pass bei Maharigaon, ca. 5000 m).

— Puthz, 1980a: 35 ( Stenus ; characters).

- DISTRIBUTION: Nepal.

subfasciatus Fairmaire, 1860b: 162 ( Stenus ; Type locality: Constantine).

— Fauvel, 1873: 77 [= 1873a: 235] ( Stenus ; characters; Algeria).

— Fauvel, 1878: 100 [= 1878a: 20] ( Stenus ; Algeria).

— Fauvel, 1886: 27 [= 1886a: 19] ( Stenus ; Algeria).

— Fauvel, 1902b: 74 ( Stenus ; Algeria; Canary Islands [Note: Specimens from Canary Islands identified as subfasciatus are misidentifications of argyrotoxus according to Puthz, 1988a: 287]).

— Bernhauer and Schubert, 1911: 167 ( Stenus ; catalog).

— L. Benick, 1929a: 19 ( Stenus ; subgenus Stenus ; characters in key).

— Scheerpeltz, 1933: 1167 ( Stenus ; subgenus Stenus ; catalog).

— Puthz, 1988a: 286 ( Stenus ; subgenus Stenus ; characters; Algeria; Morocco).

- DISTRIBUTION: Algeria, Morocco.

subfrustratus Puthz, 1968 i: 443 ( Stenus ; subgenus Hypostenus ; Type locality: Brasilien: Campobello. Also cited from Argentina).

- DISTRIBUTION: Brazil, Argentina.

subglaber Thomson, 1870 , see: glabellus Thomson, 1870 .

subgracilis L. Benick, 1941 , see: gentilis Sharp, 1886 .

subguttalis Puthz, 1969: 22 ( Stenus ; subgenus Hypostenus ; Type locality: Carin Asciuii Ghecù, 1400–1500 m, Bírmanie).

— Rougemont, 1983c: 36 ( Stenus ; characters).

— Rougemont, 1987: 707 ( Stenus ; subgenus “ Hypostenus ”; notes; Thailand).

- DISTRIBUTION: Myanmar, Thailand.

subguttatus L. Benick, 1914 , see: rufomaculatus Bernhauer, 1902 .

subhostilis Puthz, 1968: 8 ( Stenus ; subgenus Stenus ; Type locality: Mexico: Morelia).

— Puthz, 2001: 20 ( Stenus ; subgenus Stenus ; characters; Mexico).

- DISTRIBUTION: Mexico.

subimpressus Erichson, 1839 , see: pubescens Stephens, 1833 .

subimpressus Gistel, 1857 [Note: Nomen dubium], see: gistelianus E. Strand, 1917 [Note: Nomen dubium].

subiniquus Puthz, 1970: 13 ( Stenus ; subgenus Hypostenus ; Type locality: Java: Penang).

- DISTRIBUTION: Indonesia.

sublaeviventris Bernhauer, 1929, see: glacialis Heer, 1839 .

subleptocerus Puthz, 1974a: 61 ( Stenus ; subgenus Hypostenus ; Type locality: Argentina: Buenos Aires: Isla Martin Garcia).

- DISTRIBUTION: Argentina.

sublobatus Mulsant and Rey, 1861 , see: carbonarius Gyllenhal, 1827 .

submaculatus Bernhauer, 1915 h: 217 ( Stenus ; subgenus Hemistenus ; Type locality: Java: Preanger).

— Scheerpeltz, 1933: 1197 ( Stenus ; subgenus Hemistenus ; catalog).

L. Benick, 1938: 17 ( Stenus ; Java) .

— Puthz, 1970n: 307 ( Stenus ; notes).

— Puthz, 1971u: 532 ( Stenus ; subspecies of gestroi ).

— Puthz, 1981b: 146 ( Stenus ; subspecies of gestroi ; characters).

— Rougemont, 1984: 233 ( Stenus ; subspecies of gestroi ; notes; Bali; Java; Sumatra).

— Ryvkin, 1992: 45 ( Stenus ; notes; valid species).

- DISTRIBUTION: Indonesia, Philippines.

patruelis L. Benick, 1929: 94 ( Stenus ; subgenus Hemistenus ; Type locality: Binaluan, Palawan).

— Scheerpeltz, 1933: 1195 ( Stenus ; subgenus Hemistenus ; catalog).

— Puthz, 1970h: 89 ( Stenus ; subgenus Parastenus ; subspecies of submaculatus ; characters; Philippines).

drescheri L. Benick, 1938: 18 ( Stenus ; subgenus Parastenus ; Type locality: Java: G. Papandajan; G. Kawi; G. Tangkoeban Prahoe, Preanger, 4000–5000 Voet.).

— Puthz, 1970n: 307 ( Stenus ; synonym of submaculatus ).

— Rougemont, 1984: 233 ( Stenus ; synonym of gestroi submaculatus ).

submarginatus Stephens, 1833 , see: cautus Erichson, 1839 .

submarginatus Bernhauer, 1922 , see: formosanus L. Benick, 1914 .

submarginatus Bernhauer, 1912 , see: transitus L. Benick, 1921 .

submetallicus Cameron, 1930: 400 ( Stenus ; subgenus Mesostenus ; Type locality: Chakrata district: Konain. Simla Hills, alt. 7000 to 8000 feet).

— Scheerpeltz, 1933: 1197 ( Stenus ; subgenus Hemistenus ; catalog).

— Puthz, 1968j: 472 ( Stenus ; subgenus Parastenus ; characters).

— Abdullah and Qadri, 1968a: 306 ( Stenus ; characters; Pakistan).

— Scheerpeltz, 1976: 25 ( Stenus ; Nepal).

— Puthz, 1976: 7 ( Stenus ; Nepal).

— Puthz, 1986k: 962 ( Stenus ; Pakistan).

- DISTRIBUTION: Pakistan, India, Nepal.

submodicus Puthz, 1968 k: 206 ( Stenus ; subgenus Parastenus ; Type locality: Kenya:

Mt. Elgon, ö. sid., Maji ya Moto, 3710 m).

— Puthz, 1971: 331 ( Stenus ; subgenus Parastenus ; characters).

- DISTRIBUTION: Kenya.

subnitidus Bernhauer, 1939 , see: lewisius Sharp, 1874 .

subnotatus Fauvel, 1891: 95 ( Stenus ; Type locality: San Esteban; [Note: See lectotype designation by Puthz, 1969]).

— Bernhauer and Schubert, 1911: 185 ( Stenus ; subgenus Hemistenus ; catalog).

— L. Benick, 1938c: 281 ( Stenus ; characters).

— Puthz, 1969: 42 ( Stenus ; subgenus Parastenus ; lectotype designation; characters).

- DISTRIBUTION: Venezuela.

subnudus Eppelsheim, 1893 , see: alienus Sharp, 1874 .

subopacus Fauvel, 1907: 20 ( Stenus ; Type locality: Nairobi; [Note: See lectotype designation by Puthz, 1967j]. Also cited from Kilimandjaro, Kibonoto ).

— Bernhauer and Schubert, 1911: 178 ( Stenus ; subgenus Hypostenus ; catalog).

— Eichelbaum, 1913: 123 ( Stenus ; Tanzania).

— Bernhauer, 1947: 158 ( Stenus ; Deutsch-Ostafrika).

— L. Benick, 1951: 202 ( Stenus ; Tanzania).

— Puthz, 1965: 492 ( Stenus ; subgenus Hypostenus ; characters; South Africa; Tanzania; Urundi). — Puthz, 1967i: 194 ( Stenus ; characters; Kenya; Tanzania; Transvaal).

Puthz, 1967j: 12, 23 ( Stenus ; lectotype designation: Nairobi ; characters; Zaire; Kenya) .

— Puthz, 1969: 29 ( Stenus ; subgenus Hypostenus ; notes).

— Puthz, 1971: 209 ( Stenus ; subgenus Hypostenus ; Kenya).

- DISTRIBUTION: Kenya, Tanzania, Burundi, Congo, South Africa.

vanderijsti Bernhauer, 1929f: 106 ( Stenus ; subgenus Hypostenus ; Type locality: Kisantu). — Scheerpeltz, 1933: 1184 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1971: 209 ( Stenus ; synonym of subopacus alutiventris ).

alutiventris Bernhauer, 1929f: 107 ( Stenus ; subgenus Hypostenus ; Type locality: Ituri:

La Moto: Madyu).

— Scheerpeltz, 1933: 1173 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1965: 493 ( Stenus ; subgenus Hypostenus ; subspecies of subopacus ; characters).

— Puthz, 1967i: 195 ( Stenus ; subspecies of subopacus ; characters; Congo; Tanzania).

— Puthz, 1968a: 34 ( Stenus ; subgenus Hypostenus ; subspecies of subopacus ; Congo).

— Puthz, 1971: 209 ( Stenus ; subgenus Hypostenus ; subspecies of subopacus ; Zaire; Tanzania). mackayi Tottenham, 1953b: 487 ( Stenus ; subgenus Hypostenus ; Type locality: Kilimanjaro: south side, 6000–7000 ft.).

— Puthz, 1967j: 23 ( Stenus ; synonym of subopacus ).

— Puthz, 1969: 29 ( Stenus ; synonym of subopacus ).

— Puthz, 1971: 209 ( Stenus ; synonym of subopacus ).

subruandae Puthz, 1968, see: torrentum Cameron, 1951 .

subrugosus Rey, 1884 , see: providus Erichson, 1839 .

subrugosus Stephens, 1833 , see: impressus Germar, 1824 .

subsimilis L. Benick, 1942: 22 ( Stenus ; subgenus Hypostenus ; Type locality: NO-Burma: Kambaiti, 7000 ft.).

— Rougemont, 1987: 707 ( Stenus ; subgenus Hypostenus ; Thailand).

- DISTRIBUTION: Myanmar, Thailand.

subsobrinus Puthz, 1971: 260 ( Stenus ; subgenus Hypostenus ; Type locality: Guinée: Dalaba). — Puthz, 1982c: 193 ( Stenus ; notes; Fernando Poo).

- DISTRIBUTION: Guinea, Equatorial Guinea.

subthoracicus Puthz, 1970: 16 ( Stenus ; subgenus Parastenus ; Type locality: Burma:

Carin Ghecù, 1300–1400 m).

— Rougemont, 1983c: 51 ( Stenus ; characters; Thailand).

- DISTRIBUTION: Myanmar, Thailand.

subtilis Casey, 1884 , see: difficilis Casey, 1884 .

subtropicus Cameron, 1949 , see: basicornis Kraatz, 1859 .

subvastus Tottenham, 1956, see: consobrinus L. Benick, 1916 .

subviolaceus L. Benick, 1921 , see: violaceus Bernhauer, 1906 .

succinifer Rougemont, 1983c: 29 ( Stenus ; subgenus Hypostenus ; Type locality: Thailand: Chiang Mai Prov., Doi Inthanon, ca, 2000 m).

- DISTRIBUTION: Thailand.

sucinigutta Puthz, 1991d: 11 ( Stenus ; subgenus Hypostenus ; Type locality: Taiwan: [near Nanshanchi] Nantou-Hsien, 830 m).

- DISTRIBUTION: Taiwan.

sudanensis Puthz, 1965 , see: depilis L. Benick, 1951 .

sugiei Naomi, 1997c: 3 ( Stenus ; subgenus Parastenus ; Type locality: Japan: Ishikawa Pref.: Nabetani, Tatsunokuchi-cho).

- DISTRIBUTION: Japan.

sulawesicus Puthz, 1966 , see: piliferus Motschulsky, 1857 .

sulcaticeps L. Benick, 1929: 252 ( Stenus ; subgenus Hypostenus ; Type locality: Subaan, Mindoro).

— Scheerpeltz, 1933: 1183 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1991d: 23 ( Stenus ; characters).

- DISTRIBUTION: Philippines.

sulcatulus Mulsant and Rey, 1870 , see: boops Ljungh, 1810 .

sulcicollis Stephens, 1833: 295 ( Stenus ; Type locality: Suffolk; near London; Devonshire). — Stephens, 1839: 414 ( Stenus ; characters; England).

— Waterhouse and Janson, 1855: 148 ( Stenus ; characters; notes).

— Fauvel, 1869: 492 ( Stenus ; synonym of melanopus ).

— Rye, 1870a: 84 ( Stenus ; unidentifiable species).

— Bernhauer and Schubert, 1911: 185 ( Stenus ; subgenus Hemistenus ; catalog).

- DISTRIBUTION: England.

sulcifrons Eppelsheim, 1878 , see: piscator Saulcy, 1864 .

sulcipennis L. Benick, 1921b: 124 ( Stenus ; subgenus Hypostenus ; Type locality: Borneo, Sarawak).

— Scheerpeltz, 1933: 1183 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1973d: 72 ( Stenus ; characters; Borneo).

- DISTRIBUTION: Borneo.

sumbaensis Scheerpeltz, 1957c: 227 , 249 ( Stenus ; subgenus Stenus ; Type locality: Sumba: Lindiwatju).

— Puthz, 1980a: 38 ( Stenus ; characters).

- DISTRIBUTION: Indonesia.

sumbawaensis Puthz, 1969 , see: scheerpeltzianus Puthz, 1969 .

superfasciatus Puthz, 1988a: 288 ( Stenus ; subgenus Stenus ; Type locality: Algerien:

Mt. Edough).

- DISTRIBUTION: Algeria.

superlatus Puthz, 1972 , see: convergens L. Benick, 1933 .

suramensis Eppelsheim, 1880: 466 ( Stenus ; Type locality: Suramgebirge).

— Eppelsheim, 1890: 227 ( Stenus ; characters).

— Bernhauer and Schubert, 1911: 185 ( Stenus ; subgenus Hemistenus ; catalog).

— L. Benick, 1929a: 76 ( Stenus ; subgenus Parastenus ; characters in key).

— Puthz, 1972h: 125 ( Stenus ; subgenus Parastenus ; characters; Caucasus).

— Puthz, 1983e: 354 ( Stenus ; Georgia).

— Ryvkin, 1990: 203 ( Stenus ; notes; Georgia).

- DISTRIBUTION: Georgia.

suspectus Blatchley, 1910 , see: colonus Erichson, 1840 .

sutteri Scheerpeltz, 1957 , see: flavidulus Sharp, 1889 .

svanus Ryvkin, 1990: 158 ( Stenus ; subgenus Nestus ; Type locality: Gruziia, Svanetskii khr., r. Khumprer [levyi pritok r. Inguri], 1. 5 km vyshe, 20 km SV pos. Khaishi, 950 m).

- DISTRIBUTION: Georgia.

syaca Naomi, 1989e: 46 ( Stenus ; subgenus Hypostenus ; Type locality: Japan: Fukui Pref.: Ohono, Rokuroshi).

- DISTRIBUTION: Japan.

sybaris Puthz, 1971: 250 ( Stenus ; subgenus Hypostenus ; Type locality: Tanzania: Ernelo).

Hammond, 1975a: 13 ( Stenus ; notes; spelling and location of type locality is probably erroneous and is likely Ermelo , Transvaal; South Africa) .

- DISTRIBUTION: South Africa, Lesotho, Tanzania?.

pseudobconicus Scheerpeltz, 1974: 89 ( Stenus ; subgenus Hypostenus ; Type locality: Basutoland: Hensley’s Dam, 8 miles SW Leribe).

— Hammond, 1975a: 13 ( Stenus ; synonym of sybaris ).

sylvester Erichson, 1839a: 547 ( Stenus ; Type locality: Berol.; [Note: See lectotype designation by Puthz, 1967h]. Also cited from Mark Brandenburg).

— Erichson, 1840: 708 ( Stenus ; characters; Germany).

— Heer, 1841: 577 ( Stenus ; characters; Switzerland).

— Redtenbacher, 1849: 827 ( Stenus ; characters).

— Leprieur, 1851: 196 ( Stenus ; characters).

— Fairmaire and Laboulbène, 1856: 582 ( Stenus ; characters; France).

— Thomson, 1857: 226 ( Stenus ; characters; Sweden).

— Kraatz, 1857d: 766 ( Stenus ; characters; Germany).

— Redtenbacher, 1857: 226 ( Stenus ; characters; Germany).

— Thomson, 1860: 217 ( Stenus ; characters; Scandinavia).

— Rye, 1864: 61 ( Stenus ; characters; England).

— Fauvel, 1865a: 306 [= 1865b: 56] ( Stenus ; notes; France).

— Fauvel, 1873: 86 [= 1873a: 244] ( Stenus ; characters; France; Scandinavia; Germany; Switzerland; Italy).

— Fauvel, 1873b: 129 [= 1873c: 23] ( Stenus ; Russia).

— Fauvel, 1874: 331 [= 1874b: 37] ( Stenus ; characters; France).

— Redtenbacher, 1874: 246 ( Stenus ; characters).

— Seidlitz, 1875: 254 ( Stenus ; characters; Baltic region).

— Rey, 1884: 240 [= 1884a: 88] ( Stenus ; characters; France).

— Ganglbauer, 1895: 567 ( Stenus ; cited as silvester; characters; north and middle Europe).

— Reitter, 1909: 156 ( Stenus ; cited as silvester; characters; Germany).

— Bernhauer and Schubert, 1911: 167 ( Stenus ; catalog).

— Johansen, 1914: 495 ( Stenus ; subgenus Stenus ; characters; Denmark).

— L. Benick, 1929a: 25 ( Stenus ; subgenus Stenus ; characters in key).

— Portevin, 1929: 397 ( Stenus ; cited as silvester; characters; France).

— Scheerpeltz, 1933: 1167 ( Stenus ; subgenus Stenus ; catalog).

— Wüsthoff, 1934a: Tafel I ( Stenus ; illustration of aedeagus).

— Szujecki, 1961: 25 ( Stenus ; subgenus Stenus ; characters; Poland).

— Palm, 1961a: 77 ( Stenus ; subgenus Stenus ; characters; Sweden; Denmark; Norway; Finland).

— Horion, 1963: 320 ( Stenus ; subgenus Stenus ; Russia; Finland Sweden; Norway; Denmark; Baltic States; Poland; Czech Republic; Slovakia; Romania; Bosnia; Italy; France; Holland; Germany; Austria).

— Lohse, 1964: 112 ( Stenus ; subgenus Stenus ; characters; central Europe).

— Puthz, 1967h: 288 ( Stenus ; lectotype designation: Berol.).

— Puthz, 1972s: 252 ( Stenus ; Yugoslavia).

— Allen, 1978: 68 ( Stenus ; notes; may not be in Britain).

— Lucht, 1987: 96 ( Stenus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Ryvkin, 1990: 152 ( Stenus ; notes; Russia).

— Puthz, 1991f: 167 ( Stenus ; Belorussia).

— Dauphin, 1991: 285 ( Stenus ; subgenus Stenus ; characters; France).

— Hromádka, 1993: 44 ( Stenus ; subgenus Stenus ; Czech Republic; Slovakia).

— Puthz and Zanetti, 1995: 19 ( Stenus ; Italy).

— Jelínek, 1999: 155 ( Stenus ; collecting note; Czech Republic).

- DISTRIBUTION: Europe, Russia.

synonymus Gemminger and Harold, 1868 , see: humilis Erichson, 1839 .

syriacus Bernhauer, 1912 , see: ganglbaueri Bernhauer, 1905 .

syriacus Bernhauer, 1928 , see: reitterianus Bernhauer, 1928 .

syugen Namoi, 1989d: 6 ( Stenus ; subgenus Hypostenus ; Type locality: Japan: Wakayama Pref.: Mt. Gomanodan).

- DISTRIBUTION: Japan.

szechuanus Puthz, 1981d: 117 ( Stenus ; subgenus Stenus ; named for misidentification of biguttatus by Bernhauer, 1936; Type locality: China: N.O. Szechuan).

— Bernhauer, 1936: 2 ( Stenus ; cited as biguttatus [Note: Misidentification of szechuanus according to Puthz, 1981d]; China).

— Naomi, 1997f: 5 ( Stenus ; species not known in Japan).

- DISTRIBUTION: China.

szekessyi L. Benick, 1949 , see: denieri Bernhauer, 1939.

tabascensis Bernhauer, 1910b: 366 ( Stenus ; Type locality: Mexiko: Teapa , Tabasco).

— Bernhauer and Schubert, 1911: 178 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1974a: 68 ( Stenus ; characters).

- DISTRIBUTION: Mexico.

tacitus Puthz, 1967 k: 10 ( Stenus ; subgenus Hypostenus ; Type locality: N. Rhodesia: Abercorn, 1800 m, galerie forestière de la Mwenga).

— Puthz, 1971: 272 ( Stenus ; subgenus Hypostenus ; characters; Zambia; Zaire).

— Puthz, 1972a: 5 ( Stenus ; characters; Zaire).

- DISTRIBUTION: Zambia, Congo.

tacomae Casey, 1884 , see: bilineatus J. Sahlberg, 1871 .

tahoensis Casey, 1884: 139 ( Stenus ; Type locality: Lake Tahoe , Nevada).

— Bernhauer and Schubert, 1911: 171 ( Stenus ; subgenus Tesnus ; catalog).

— Puthz, 1980k: 35 ( Stenus ; notes).

— Campbell and Davies, 1991: 112 ( Stenus ; Canada).

- DISTRIBUTION: USA, Canada.

pugetensis Casey, 1884: 140 ( Stenus ; Type locality: Thurston County, Washington Territory ).

— Bernhauer and Schubert, 1911: 171 ( Stenus ; subgenus Tesnus ; catalog).

— Sanderson, 1957: 268 ( Stenus ; subgenus Tesnus ; characters; Washington; Oregon).

— Puthz, 1980k: 35 ( Stenus ; synonym of tahoensis ).

vacuus Casey, 1884: 141 ( Stenus ; Type locality: San Mateo , California).

— Bernhauer and Schubert, 1911: 171 ( Stenus ; subgenus Tesnus ; catalog).

— L. Benick, 1938c: 268 ( Stenus ; characters).

— Puthz, 1980k: 35 ( Stenus ; synonym of tahoensis ).

taigae Poppius, 1909: 11 ( Stenus ; Type locality: Taiga-See bei Shigansk).

— Bernhauer and Schubert, 1911: 168 ( Stenus ; catalog).

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Genus

Tesnus

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Genus

Edaphus

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