Drasteria flexuosa flexuosa ( Ménétriés, 1849 ), flexuosa (Menetries, 1849

Yu, Alexey & Korb, Stanislav K., 2019, A revision of the genus Drasteria of Central Asia and Kazakhstan with special attention to the adjacent areas (Lepidoptera: Erebidae), Zootaxa 4673 (1), pp. 1-104 : 66

Drasteria flexuosa flexuosa ( Ménétriés, 1849)

(= D. flexuosa caspica ( Staudinger, 1901) , = D. flexuosa singularis ( Kollar, 1849)) ( Figs. 175–177 , 190–192 (imago), 258, 259 (male genitalia), 290 (female genitalia))

= singularis Kollar, 1849: 53 ( Ophiusa Singularis ). Type locality: “Südpersien” (by the original description).

Type material. Not found; probably, lost.

= caspica Staudinger in Staudinger & Rebel, 1901: 244 ( Leucanitis Flexuosa Mén. v. Caspica). Type locality: “Lac. Casp. lit. (Ross. m. or. et Pers. s.)” (by the original description); “Birutschi” (Biruchya Kosa peninsula, 45°43′04″N, 47°34′45″E) (by the lectotype designation, designated here).

Type material. Lectotype male (designated here in accordance with ICZN Art. 74) ( Figs. 187–189 ), labelled: “ex coll. 1/4 | Staudinger” (white paper, printed with handwritten numbers), “Origin.” (pink paper, printed), “Birutschi | Caspi-Meer” (green paper, handwritten), “ LECTOTYPUS | Leucanitis flexuosa var. cas- | pica Staudinger, Rebel, 1901 | Cat. Lep. paraearct. Faunengeb. 1: 244 | Korb, Matov, Volynkin des. | 2016, Feb. 1–4 ” (red paper, printed) ( ZMHU) . Paralectotypes: 3 males ( ZMHU) .

Erschov, 1874: 56 [Turkestan city]; Butler, 1892: 316 [Abscheron]; Staudinger, Rebel, 1901: 244 [Ili]; Meinhard, 1908: 26 [Dzharkent]; John, 1910: 617 [Kizil-kum, Perovsk, Dshulek,Aralsee]; Warren 1913:389 [Ili]; Wiltschire, 1957 [widespread in Iraq]; Kuznetzov, 1960: 87 [Askhabad]; Daricheva, 1965: 57 [Askhabad; Bairam-Ali; Tashkepri; Akar-Cheshme; Kushka; Kara-Kala; Chardzhou]; Stshetkin, 1965: 168 [Buritau Mts., Sand Pass, Kolkhozabad, Dzhilikul,Aruktau Mts., Khalka- Kul lake, Darya-Kul lake]; Sviridov, 1971: 47 [Kushka, Akar-Cheshme]; Shek, 1972: 143 [Chimkent Prov.]; Sukhareva, 1972: 62 [Ayakguzhumdy, Zhamansai, Shafrikan]; Aibasov, 1975: 122 [Irgiz, Malye and Bolshie Barsuki, Altynchokusu plateau]; Shek, 1975: 415 [Guryev, Alma-Ata, Taldy-Kurgan Prov]; Daricheva et al., 1983: 38 [Lambe; Deinau]; Radzhabova, 1985: 25 [Kuraminsky Mts.: Tashbulak valley, Cholata]; Radzhabova, 1992: 181 [Uchkhoz, Sovkhoz “Syrdarya”, Kolkhoz imeni Frunze, Katarbulak, Cholata, Kok-Kurtak, Zafarabad]; Weisert, 1997: 73 [Provinz Mary, Karakum-Wiiste, 12km N Zakhmet; Provinz Mary, 14km N Zakhmet; Provinz Chardzhou, Suit Kugitangtau, Gaynar-baba-See; Usbekistan, N-Kugitangtau, Leilakhansei]; Lehmann et al., 1998: 526 [33.37’N, 79.21’O, Sarytogai, Tal des Flusses Sharyn, ca 20 km vor der Mündung in den Ili, 666m; Ili area, Charyn river, 400m; Alma-Ata area, Ili river, Taschkarasu, 450m]; Poltavsky et al., 1998: 108 [Bairam-Ali, Lambe, Deinau, Tschardzhou, Repetek, Mary, Badkhyz, Kyzyl-Arvat, Kara-Takyr, Kara-Kala, Yaskhan, Danata, Dardja, Bugdaili]; Ivinskis, Miatleuski, 1999: 203 [Ashkhabad, Sandykatschi, Kov-Ata]; Anikin et al., 2000: 332 [Astrakhan]; Hacker, Miatleuski, 2001: 814 [Sajkhin; Urda]; Poltavsky, Ilyina, 2002: 17 [Ribatchij, Gunib]; Poltavsky, Nekrasov, 2002: 26 [Kalmykia Republic, Daghestan Republic]; Goater et al., 2003: 75 [South Russia: Naryn area, southern Kirgisian steppe, Kazakhstan: Sajkhin and Urda; outside Europe from Egypt through Turkey, Caucasus, Asia Minor, Afghanistan to Mongolia): 254 (Alma Ata Gebiet, Taschkarasu, 500 m]; Poltavsky, Ilyina, 2003: 171 [plateau Gunib]; Benedek, Bálint, 2009: 162 [Prov. Almaty, Taukum desert, 10 km W of Usharal, 450 m, N44°12’ E076°47’]; Poltavsky et al., 2010: 96 [Botkul]; Gorbunov, 2011: 57 [deserts of West Kazakhstan]; pl. 8, fig. 7 [Mamekkazgan]; Gorbunov et al., 2011: 76 [Ustyurt Nature Reserve, Bostankum sands, 40 km NE Zhanaozen, 43°39‘ N, 53°08‘ E; Ustyurt Nature reserve, Besokty dwell, 18 km E Senek, 43°20‘ N, 53°36’ E; Northern slope of Ustyurt Nature reserve, 43°24’ N, 54°33’ E; Ustyurt Nature Reserve, S of Karashek Mt. 18 km N of Akkuduk, 43°06‘54“ N, 54°11‘40“ E; Ustyurt Nature reserve, Kertaktay, 43°17’ N, 54°27‘ E; Ustyurt Nature reserve, Kendyrli dwell, 42°57’ N, 54°41‘ E; Ustyurt Nature reserve, Zhomarta, 13 km SE Akkuduk, 42°55’ N, 54°15’ E; Ustyurt Nature reserve, Onere dwell, 42°36’ N, 54°08‘ E]; Korb, 2013: 28 [Saksaul forest 56.5 km S of Kugaly, Altyn Emel Nature Reserve, N 43 57.688 E 78 46.164, 645 m]; Shovkoon, Trofimova, 2016: 607 [West Kazakhstan Prov., N 48°38‘; E 50°41‘; Atyrau Prov., N 48°28‘; E 51°18‘; Atyrau Prov., 15 km E Komsomolsky village, N 47°19‘, E 53°50‘; Kyzyl-Orda Prov., Karatup penin, Aral lake, N 46°20‘; E 59°43‘; Mangistau Prov., Tuyb-Karagan penin. Caspian sea, N 44°30‘; E 51°01‘; Mangistau Prov., Aktau Mts., N 44°16‘; E 52°10‘; Mangistau Prov., Tuyb-Karagan penin., N 44°14‘; E 50°47‘; Mangystau Prov., N 44°07‘; E 51°50‘].

Habitat ( Figs. 300, 302, 303 ). Deserts and semideserts. Elevation: 200– 700 m.

Distribution. Deserts of Central Asia, South Russia and Kazakhstan, Turkey, Israel and Iran.

Material examined (in total: 544 specimens). Afghanistan. 1 ♂, 14.05.1911, Faisabad , [leg. Zarudny], ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♂, 12.06.1970, Prov. Samangan, obere Tang-e-Tashawighan , 500 m, leg. Naumann ( HNHM) . Armenia. 1 ♂, 2 ♀, 12– 16.05.1925, pr. Eriwan, Parakar , leg. A. Schelkovnikov ( ZISP) ; 1 ♀, 4.11.1967, Aivopat , leg. E. Miljanovskij ( ZISP) ; 1 ♀, 12.07.1958, Norashen , leg. unknown ( ZISP) . Azerbaijan. 1 ♀, Ordubad, ex coll. Tshetverikov ( ZISP) ; 2 ♂, 1 ♀, 5.05.[18]83, Ordub [ad], leg. Chr [istoph], ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 2 ♀, Baku, leg. А. Tebel, ex coll. Acad. Petrop. ( ZISP) ; 1 ♂, 1 ♀, 6– 10.07.1906, Baku, leg. L. Bianki ( ZISP) ; 1 ♀, 24.07.1933, Disar near Ordubad, leg. Znoiko ( ZISP) ; 1 ♀, 11.06.1937, Dzhuga , [leg. M. Ryabov] ( ZISP) ; 1 ♂, 22.06.1979, Uch-Topa , leg. A. Danchenko, ex coll. A. Nekrasov ( ZISP) . China. 1 ♂, 15.05.1879, Tian Chian , leg. Alph [eraky], ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♀, 1.07.1884, Huan- Djin , leg. Potanin, ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♀, 2.05.1909, Kashgaria, Yaz-Bulak , leg. Divnogorsky ( ZISP) ; 1 ♂, 12.05.1909, Kashgaria, Yangiabad , leg. Divnogorsky, ex coll. Tshetverikov ( ZISP) . Georgia. 1 ♂, 10.08.1928, flum. Kura, u. Achalzich , 10 km Or. dist., leg. A. Schorygin ( ZISP) . Iran. 1 ♂, 4 ♀, 16.0 5– 5.06.1955, Beludzhistan , Zakhedan, leg. Schteinberg ( ZISP) ; 1 ♂, 18.05.1955, Beludzhistan, Khash , leg. Schteinberg ( ZISP) ; 1 ♀, 11.10.1955, Beludzhistan, Bampur , leg. Stsherbinovsky ( ZISP) ; 1 ♂, 28– 30.05.1963, Derbend , 25 km N v. Teheran, 2000 m, leg. Kasy and Vartian ( HMNH) ; 1 ♂, 04.2008, Prov. Kerman, Qohrud Mts., 2000 m, Küh-e-Madvar , leg. T. Hácz ( NHNM) . Israel. 1 ♂, 1 ♀, Jordanthal , leg. M. Bartel, ex coll. O. John ( ZISP) . Kazakhstan. 1 ♂, 1 ♀, 7.06.[?], Uzun-Ata valley , leg. Zarudny ( ZISP) ; 1 ♂, 2 ♀, Sir [Syr Daria], [leg. Eversmann], ex coll. Eversmann ( ZISP) ; 3 ♂, 3 ♀, 05– 2.08.1903, Timur st. , 50 km from Turkestan, leg. Klare ( ZISP) ; 1 ♀, 15.05.1905, Perovsk , leg. Shell’, ex coll. Mus. Acad. Petrop. ( ZISP) ; 3 ♀, 18– 23.05.1905, Dzhulek , [leg. Shell’], ex coll. Mus. Acad. Petrop. ( ZISP) ; 1 ♀, 17.05.1907, prope Dzharkent , ad fl. Ussek, M. Divnogorsky ( ZISP) ; 1 ♀, 19.05.1907, Kasalinsk , leg. Krainev ( ZISP) ; 1 ♂, 17.06.1907, Inderskischer Salzsee , leg. M. Bartel ( ZISP) ; 3 ♀, 1908, Syr-Darja, Perowsk , leg. Nikolsky ( ZISP) ; 1 ♂, 1908, Baigacum , leg. Androsow, ex coll. O. John ( ZISP) ; 1 ♂, 1 ♀, 9– 10.05.1908, Syr-Darja, St. Baigakum , leg. Malyschew, ex coll. O. John ( ZISP) ; 3 ♂, 1 ♀, 26.04.1909, 10– 24.07.1910, Syr-Darja, Dshulek , leg. Koshantschikoff, ex coll. O. John ( ZISP) ; 1 ♂, 1.05.1909, Perovsk , leg. Е. Miller ( ZISP) ; 5 ♂, 10 ♀, 1.05– 24.07.1909, Syr-Darja, Aj-Darle , leg. Koshantschikoff, ex coll. O. John ( ZISP) ; 1 ♂, 1 ♀, 16– 19.05.1910, prov. Syr-Darja, deser. Mujun-Kum , lac. Kargaly-Kul, leg. A. Golbeck ( ZISP) ; 6 ♂, 9 ♀, 5.07– 7.08.1911, Syr-Daria, Baigacum , leg. Koshantschikoff, ex coll. O. John ( ZISP) ; 6 ♂, 2 ♀, 1915, Semiretshye Prov., Dzharkent , [leg. Dyakonov], ex coll. Dyakonov ( ZISP) ; 4 ♂, 5 ♀, 04.1915, Dzhar-Kuduk on the S of Dzhar- kent, leg. V. Kavrigin ( ZISP) ; 12 ♂, 11 ♀, 06.1915, Chulokai S of Dzharkent, leg. V. Kavrigin ( ZISP) ; 1 ♂, 14.07.1930, Kzyl-Orda , leg. unknown ( ZISP) ; 1 ♀, 30.05.1932, Aralsk , leg. unknown ( ZISP) ; 2 ♂, 1 ♀, 4.07.1940, Aral distr., Lai-Kul lake , leg. E. Shumakov ( ZISP) ; 4 ♂, 7 ♀, 8.07.1948, 17– 18.07.1950, 15.08.1952, 7.07.1953, Dzhusaly, leg. Bundel ( ZISP) ; 1 ♀, 14.07.1952, Syr-Darya, Dzhologosh , leg. Bundel ( ZISP) ; 1 ♂, 1 ♀, 9.07.1954, Turkestan st. , leg. A. Bundel ( ZISP) ; 1 ♂, 23.08.1955, Mangyshlak, Saubei spurs, leg. Grunin ( ZISP) ; 3 ♂, 2 ♀, 26.07– 11.08.1955, Mangyshlak, Beke , leg. Grunin ( ZISP) ; 1 ♂, 6.06.1966, 40 km W Kyzyl-Orda , leg. Pastukhov ( ZISP) ; 1 ♂, 7.06.1966, near Tartugai , leg. Pastukhov ( ZISP) ; 2 ♂, 3.08.1982, Barsa-Kelmes Nature Reserve , [leg. D. Piryulin] ( ZISP) ; 1 ♀, 2.05.1990, Zhel-Turanga , leg. S. Toropov, ex coll. A. Nekrasov ( ZISP) ; 2 ♀, 29.05.1966, 80 km NW Turkestan, Akkum , leg. Pastukhov ( ZISP) ; 1 ♂, 28.08.1986, Almaata Prov., Karaoi , leg. V. Mironov ( ZISP) ; 1 ♂, 8.05. 1990, 130 km NE Alma-Ata, right shore of Ili river, Mynbulak valley , leg. Amerguzhin ( ZISP) ; 1 ♂, 10.07.2016, Ili river valley, Akzhar village , at light, 44º52’ N 75º53’ E, leg. P. Gorbunov ( ZISP) ; 1 ♂, 21.05.1930, Syrdaryinsky Karatau Mts., Turkestan vic., leg. A.V. Zvetaev ( ZMMU) ; 1 ♀, 1951, Syrdaryinsky Karatau Mts., Mynzhilki Mt. , leg. A.V. Zvetaev ( ZMMU) ; 1 ♂, 1.05.2011, between Katutau Mts. and Singing Dunes in Altyn- Emel Nature Reserve , 400 m, leg. P.Egorov ( SKK) ; 2 ♀, 30.06– 1.07.2012, Almaty Prov., Panfilov distr., 25 km SW of Koktal, Ili river , leg. S. Rybalkin ( CKO) ; 1 ♂, 1 ♀, 6– 7.05.2013, Almaty Prov., 17 km SSW of Topar , sands, leg. S.A. Knyazev ( CKO) ; 1 ♂, 7– 8.05.2013, 28 km NE of Bakanas, Ili river , leg. S.A. Knyazev ( CKO) ; 1 ♂, 1– 3.07.2015, Ili river valley near Koktal bridge, 43°57.818’N, 79°36.322’E, 500m, leg. S.K. Korb ( SKK) ; 1 ♂, 5– 6.05.2017, Kyzylorda Prov. , 15 km NE of Kyzylorda, sands, leg. S.A. Knyazev ( CKO) . Kyrgyzstan. 1 ♂, 6.05.[?], Toguz Bulak , leg. Zarudny ( ZISP) ; 1 ♂, 31.05.1910, Mil-Buluk , leg. Zarudny ( ZISP) ; 1 ♀, Issyk Kul , leg. M. Bartel, ex coll. O. John ( ZISP) . Russia. 1 ♀, Volgograd region: Sarep [ta], leg. unknown ( ZISP) ; 1 ♂, Astrakhan region: 60 verst from Astrakhan, Biruchaya Kosa , leg. Alph [eraky], ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♀, 26.07.1910, Stavropol area: Atshykulak , [leg. I. Filipyev], ex coll. N. Filipyev ( ZISP) ; 4 ♂, 5 ♀, 4.06– 4.07.1911, Kuma upper course, Zimnyaya Stavka, leg. Uvarov ( ZISP) ; 1 ♀, 28.07.1967, Astrakhan Prov., salted land, leg. Pen- chukovskaya ( ZISP) ; 3 ♂, 4.06.1970, Marfino , leg. A. Lvovsky ( ZISP) ; 1 ♂, 1 ♀, 12– 15.06.1970, Kurchenko , leg. A. Lvovsky ( ZISP) ; 1 ♀, 25.06.1970, Dosang , leg. A. Lvovsky ( ZISP) ; 1 ♀, 14.08.2002, Republic of Kalmykia: Chernozemelsky distr., Artezian , leg. O. Saranova ( ZISP) ; 1 ♂, 30.04.2010, Astrakhan Prov., Dosang , leg. S.K. Korb ( SKK) ; 2 ♂, 12– 14.06.2018, Aidarly env., 43.993779 N, 79.483105 E, leg. A. Litovtsev ( SKK) . Tajikistan. 1 ♂, 26.05.[18]85, Kulyab , leg. unknown, ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♂, 2 ♀, 12.06.1910, Kabadian , leg. Zarudny ( ZISP) ; 1 ♀, 5.06.1945, Stalinabad , leg. Gussakovsky ( ZISP) ; 1 ♀, 17.06.1963, N Tajiki- stan, leg. Kamenkova ( ZISP) ; 1 ♂, 1 ♀, 13.04.1966, Gissarsky Mts., Dzhar-Kurgan , leg. Danilevsky ( ZISP) ; 1 ♂, 11.07.1990, Pamir Mts., Dusty , 2100 m, leg. V. Lukhtanov ( HNHM) ; 1 ♂, 1 ♀, 23.04.2010, 8.07.2010, SW Tajiki- stan, 15 km SE Shaartuz, sands, 300 m, between Dzharkurgan and Kurdzhalol , leg. O. Pak ( OPB) ; 1 ♂, 12– 13.06.2012, Prov. Khalton, distr. Kulab, near Vose , mts. Khodzhamumin , 1200 m, leg. E. Rutjan ( OPB) . Turkey. 1 ♂, 28.04– 1.05.1989, Prov. Urfa, Halfeti, valley of Euphrat , 500 m, 37º52,5´E, 37º14,5´N, leg. Fábián, Ronkay and Ronkay ( HNHM) . Turkmenistan. 1 ♀, Cheliken , leg. unknown, ex coll. Tshetverikov ( ZISP) ; 1 ♂, 23–25.04.[?], Chardzhui , leg. Zarudny ( ZISP) ; 3 ♀, 28.04.[?], 135 verst from Chardzhui upper Amu-Darya river, leg. Zarudny ( ZISP) ; 1 ♂, 28.04.[?], Sary-Yazy , leg. Lekarev, ex coll. Nekrasov, slide N 2438 Nekrasov ( ZISP) ; 1 ♂, 7.05.[?], Kelif , leg. unknown ( ZISP) ; 3 ♂, 3 ♀, 7–18.05.[18]82, 05.[18]84, Askhabat , leg. Chr [istoph], ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♂, 1 ♀, 6–8.04.[18]92, Tedshen , [leg. Christoph], ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♂, 12.05.[18]94, Kazandzhin , leg. unknown, ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♂, 5.05.[18]95, Tedzhen river, Akar-Cheshme , leg. Korzhinsk. ( ZISP) ; 1 ♀, 18.04.1906, Askhabad , [leg. Meinhardt], ex coll. Meinhardt ( ZISP) ; 2 ♀, 28.04.1906, Transcaspian Prov., Tash-kipri , leg. Demokidov ( ZISP) ; 1 ♂, 29.04.1906, Kushka , leg. Demokidov ( ZISP) ; 1 ♀, 12.06.1906, Bairam-Ali , leg. Demokidov ( ZISP) ; 2 ♂, 6 ♀, 1– 7.05.1909, Murgabskoe Gosudarevo Imenie , leg. K. Demokidov ( ZISP) ; 1 ♂, 13.06.1910, Bairam-Ali , leg. Uva- rov ( ZISP) ; 6 ♂, 8 ♀, 20.05– 14.09.1913, 10.05.1915, Chardzhui on Amu-Darya river, leg. Samokish ( ZISP) ; 2 ♂, 3 ♀, 10.07– 26.08.1926, Iolotan , leg. V. Kuzericky ( ZISP) ; 1 ♀, 1927, near Mary, leg. P.Donov ( ZMMU) ; 1 ♀, 27.04.1927, Buchara occ., loc. Tshardzhui , leg. K. Malyshev ( ZISP) ; 4 ♂, 9 ♀, 20.05– 9.09.1930, Bairam-Ali , leg. Bogush, slide N 6125 Ryabov ( ZISP) ; 1 ♂, 1 ♀, 18.05.1934, Samsonovo , leg. E. Luppova ( ZISP) ; 1 ♂, 3 ♀, 21– 25.05.1934, left shore of Amu-Darya river near Kerka, Khalach, leg. E. Luppova ; 1 ♀, 27.05.1934, Khalag-Kerkim , leg. E. Luppova ( ZISP) ; 1 ♂, 1 ♀, 4–5.1937, Repetek , leg. Lavrov ( ZISP) ; 1 ♂, 11.04.1946, Bairam-Ali vicinity, leg. A.V. Zvetaev ( ZMMU) ; 1 ♀, 24.04.1951, Top-Yatan lake, Uzboi , leg. Schteinberg ( ZISP) ; 8 ♂, 3 ♀, 20.05.1952, 17.05– 3.07.1953, 40 km N Kyzyl-Arvat, Kara-Bogaz , leg. Maslennikova ( ZISP) ; 2 ♂, 10– 19.06.1952, Repetek , leg. Goryshina ( ZISP) ; 1 ♀, 22.08.1952, Askhabad Prov., Kara-Kala , VIR garden, leg. V. Kusnetzov ( ZISP) ; 1 ♂, 30.04.1953, 40 km N Kyzyl-Arvat, Kara-Bogaz , leg. V. Kusnetzov ( ZISP) ; 1 ♂, 20.05.1953, 15 km N Kizil-Arvat, Takyrabad , leg. Maslennikova ( ZISP) ; 3 ♂, 2 ♀, 4– 6.06.1953, Kara-Kum desert, Shasenem , leg. Kiryakova ( ZISP) ; 1 ♂, 1 ♀, 4– 9.07.1953, Kelif , leg. A. Bundel ( ZISP) ; 5 ♂, 9 ♀, 12.07– 22.08.1953, 4.06– 10.07.1954, 13.05– 22.06.1955, Askhabad circ., leg. W. Potopolskij ( ZISP) ; 1 ♂, 4 ♀, 14.06– 15.08.1962, Tedzhen , sovkhoz 40 let VLKSM, leg. Gullyev ( ZISP) ; 4 ♀, 11.05.1965, 12– 14.05.1966, Repetek , leg. A. Tsvetaev, ex coll. A. Nekrasova ( ZISP) ; 1 ♂, 24.09.1965, Repetek , leg. A.V. Zvetaev ( ZMMU) ; 2 ♂, 1 ♀, 3.05– 15.06.1971, Murgab distr. , kolkhoz Tazemurdush, leg. Kamalov ( ZISP) ; 1 ♀, 30.04.1979, Chardzhou , leg. unknown, ex coll. A. Nekrasov ( ZISP) ; 1 ♀, 4.05.1979, Badkhyz, Kyzyldzhar , leg. Reznik ( ZISP) ; 1 ♂, 1.06.1980, Krasnovodsk , leg. P. Kazaryan, ex coll. A Nekrasov ( ZISP) ; 1 ♀, 6.10.1985, 5 km N Askhabad , leg. M. Falkovich ( ZISP) ; 1 ♂, 17.04.1987, Geok-Tepe distr., Babarab , leg. L. Petrikevich ( ZISP) ; 1 ♀, 17.04.1988, Geok-Tepe distr., Babarab , leg. M. Prokofiev ( ZISP) ; 1 ♀, 26.08.1990, Mary, kolkhoz Vatai , leg. V. Mironov ( ZISP) ; 1 ♀, 31.07.1991, Bairam-Ali, Karakumkanal sovkhoz, leg. V. Mironov ( ZISP) . Uzbekistan. 1 ♀, Bokara [Bukhara], ex coll. Acad. Petrop. (holotype) ( ZISP) ; 1 ♀, 29– 30.05.[?], Termez, leg. unknown ( ZISP) ; 1 ♂, 5 ♀, 1895, Kara-Tyube , leg. Ellik ( ZISP) ; 2 ♀, Russky Turkestan , [leg. Fedchenko], ex coll. Erschov ( ZISP) ; 1 ♀, Turkestan , ex coll. Erschov ( ZISP) ; 1 ♂, 1 ♀, 3– 10.05.1892, Samar- kand, leg. O. Herz, ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 2 ♂, 05.1903, border between Kyzyl-Kum desert and Golodnaya Step near Amu-Darya river ., leg. G. Yakobson ( ZISP) ; 1 ♂, 10.05.1903, Kyzyl-Kul lake near Golodnaya Step , leg. N. Ivanov ( ZISP) ; 1 ♂, 3 ♀, 6– 30.05.1903, Golodnaya Step in Khodzhent Prov. , leg. G. Yako- bson ( ZISP) ; 1 ♀, 1911, Bukhara , leg. A. Golbeck ( ZISP) ; 2 ♀, 9– 16.08.1908, prov, Samarkand, stac. Obrutshovo , leg. A. Golbeck ( ZISP) ; 1 ♀, 20.08.1910, Karakalpakia, Khodzheili , leg E. Fisher ( ZISP) ; 1 ♂, 3 ♀, 19.04– 13.09.1911, Buchara, stc. Farab , leg. A. Golbeck ( ZISP) ; 2 ♂, 4.05.1911, Buchara occ., ad lac Dengiz-Kul, kischlak Assia , leg. unknown, ex coll. Tshetverikov ( ZISP) ; 1 ♀, 11.06.1911, Tashkent , leg. unknown ( ZISP) ; 1 ♂, 14.06.1911, Kran- tauli, leg. Molchanov ( ZISP) ; 1 ♂, 1 ♀, 17.06.1911, Amu-Darya river valley, Kubetau Mt. , leg. Molchanov ( ZISP) ; 1 ♂, 30.06.1911, Tashkent distr. , leg. unknown ( ZISP) ; 1 ♀, 16.07.1911, Kuvapsh, Dzherma , leg. Molchanov ( ZISP) ; 1 ♀, 26.07.1911, Nukus , leg. Molchanov ( ZISP) ; 5 ♂, 3 ♀, 21.04– 20.07.1912, 21.04.1913, Termez , leg. Kiritshen- ko, ex coll. Dyakonov ( ZISP) ; 1 ♀, 14.05.1914, Samarkand , leg. unknown ( ZISP) ; 2 ♂, 1 ♀, 13.05.1926, 18.06.1927, Bukhara , leg. Yakhontov ( ZISP) ; 1 ♂, 1.06.1926, Bukhara, Guzars. , leg. A. Gerasimov ( ZISP) ; 1 ♂, 1 ♀, 3.05.1927, Khiva, Ravat , leg. L. Zimin ( ZISP) ; 2 ♂, 3 ♀, 23.05– 3.09.1927, Bukhara , leg. V. Zolotov ( ZISP) ; 1 ♀, 2.08.1927, Khiva, Ravat , leg. V. Gussakovsky ( ZISP) ; 2 ♂, 6– 22.04.1928, Buchara , leg. A. Gerasimov ( ZISP) ; 3 ♂, 1 ♀, 20.04– 14.08.1928, Buchara, Baga-Absal , leg. A. Gerasimov ( ZISP) ; 1 ♀, 17.05.1928, Buchara, Kujumasar , leg. A. Gerasimov ( ZISP) ; 3 ♀, 22– 30.05.1928, Buchara, Chargush , leg. A. Gerasimov ( ZISP) ; 1 ♂, 11.06.1928, Samar- kand Prov., Khargush , leg. Gerasimov ( ZISP) ; 1 ♂, 12.07.1928, Buchara, Kara-Kul , leg. A. Gerasimov ( ZISP) ; 1 ♀, 8.05.1929, Buchara, Kisil-Tepe , leg. A. Gerasimov ( ZISP) ; 2 ♂, 6.05– 22.06.1930, Samarkand Prov., Khatyrginsky distr. , Changir, leg. Zimin ( ZISP) ; 1 ♂, 12.05.1931, Kammashi , leg. Rodd ( ZISP) ; 3 ♂, 16.05– 5.06.1931, 21.06.1932, Kammashi N-O Guzar , leg. Gussakovsky ( ZISP) ; 5 ♂, 10 ♀, 4.08.1938, Ferghansky Mts., Kokand, Buvaida , leg. A.V. Zvetaev ( ZMMU) ; 1 ♀, 14.07.1957, Ursatyevskaya , leg. A. Bundel ( ZISP) ; 1 ♀, 19.07.1958, Fergana Prov., Kokand, Buvaida , leg. A. Tsvetaev, ex coll. A. Nekrasov ( ZISP) ; 1 ♂, 2 ♀, 13.04.1959, Dzhar-Kurgan , leg. A.V. Zvetaev ( ZMMU) ; 2 ♀, 19– 25.04.1965, Kyzyl-Kum desert, Ayak-Agytma , leg. Pastukhov ( ZISP) ; 8 ♂, 9 ♀, 13– 27.04.1965, 40 km E Dzhingildy, Ayakguzhumdy, leg. Pastukhov ( ZISP) ; 20 ♂, 26 ♀, 28.04– 17.06.1965, 29.04– 30.05.1966, 11.05– 20.10.1967, 11– 18.09.1968, 12.05– 5.09.1969, 2.04– 18.08.1970, 14.09– 27.10.1971, 19.04– 16.06.1975, 29.04.1976, 40 km E. Dzhingildy, Ayakguzhumdy, leg. M. Falkovich ( ZISP) ; 1 ♀, 2.05.1965, Kyzylkum desert, 90 km S Tamdy-Bulak, Aznek , leg. Pastukhov ( ZISP) ; 1 ♂, 13.05.1965, Kyzylkum, Bukantau Mts. , leg. Falkovich ( ZISP) ; 7 ♀, 26.05– 12.06.1965, 40 km E Dzhingildy, Ayakguzhumdy , leg. Zabello ( ZISP) ; 1 ♀, 20.06.1965, Bukhara , leg. Falkovich ( ZISP) ; 5 ♂, 2 ♀, 14– 21.04.1966, 40 km E Dzhingildy, Ayakguzhumdy , leg. I. Sukhareva ( ZISP) ; 1 ♂, 2 ♀, 17.04– 12.05.1966, 140 km NW Shafrikan, Zhamansai , leg. I. Sukhareva ( ZISP) ; 1 ♂, 1 ♀, 23– 24.04.1966, Kyzylkum, Zhamansai , leg. M. Falkovich ( ZISP) ; 1 ♂, 7.05.1966, Kyzylkum desert, 60 km SO Uchkuduk, leg. Pastukhov ( ZISP) ; 1 ♂, 1 ♀, 25.05.2010, Kyzylkum, no collector data ( CKO) .

Figures

FIGURES 166–186. Drasteria specimens. 166–171. D. tenera (Staudinger, 1877), lectotype (166: upperside; 167: underside; 168: labels), Koktal bridge, Kazakhstan (169), Panfilov, Kazakhstan (170), Dubin environs near Ili river, Kazakhstan (171). 172–174. D. antiqua (Staudinger, 1889), lectotype (172: upperside; 173: underside; 174: labels). 175–177. D. flexuosa flexuosa (Ménétriés, 1847), lectotype (175: upperside; 176: underside; 177: labels). 178–186. D. flexuosa mongolica (Staudinger, 1896), lectotype (178: upperside; 179: underside; 180: labels), holotype of pulverosa Wiltshire, 1969 (181: upperside; 182: underside; 183: labels), holotype of intermedia Ronkay, 1985, holotype (184: upperside; 185: underside; 186: labels).

FIGURES 187–207. Drasteria specimens. 187–189. D. flexuosa caspica (Staudinger, 1901), lectotype (187: upperside; 188: underside; 189: labels). 190–192. D. flexuosa (Ménétriés, 1847), Topar, Kazakhstan (190), Koktal bridge, Kazakhstan (191), Aksuat, Kazakhstan (192). 193–198. D. sinuosa (Staudinger, 1884), lectotype (193: upperside; 194: underside; 195: labels), Jussufabad, Iran (196–198). 199–201. Drasteria yerburyi (Butler, 1892), lectotype of albofasciata John, 1917 (199: upperside; 200: underside; 201: labels). 202–207. D. mongoliensis Wiltshire, 1969, holotype (202: upperside; 203: labels), paratype (204), Bulagan Aimak, Mongolia (205–207).

FIGURES 297–304. Drasteria habitats. 297. 25 km N of Panfilov, Dzhungarian Alatau, Kazakhstan (one of the authors in the collecting trip 2010). 298. Ili river valley near Koktal bridge, Almaty Prov., Kazakhstan (Russian Entomological Society and Institute of Zoology of Kazakhstan collecting trip, 2010). 299. Sands near Dosang environs, Astrakhan Prov., Russia. 300. Saksaul forest near Bakanas, Almaty Prov., Kazakhstan. 301. Chulak gorge, foothills of Dzhungarian Alatau Mts. in the Altyn-Emel Nature Reserve, Kazakhstan. 302. Ili river shore near Singing Dunes, Altyn-Emel Nature Reserve, Kazakhstan. 303. Katutau mountains, Altyn-Emel Nature Reserve, Kazakhstan. 304. Koram environs, northern slopes of Transilian Alatau Mts., Kazakhstan. Photos: S.K. Korb.