Haplodrassus triangularis, Bosmans & Kherbouche-Abrous & Benhalima & Hervé, 2018

Bosmans, Robert, Kherbouche-Abrous, Ourida, Benhalima, Souâd & Hervé, Christophe, 2018, The genus Haplodrassus Chamberlin, 1922 in the Mediterranean and the Maghreb in particular (Araneae: Gnaphosidae), Zootaxa 4451 (1), pp. 1-67 : 7-8

Key to the Haplodrassus species of the Maghreb region

MALES

1 Cymbium with abundant, strong spines (fig. 71)................................................... nigroscriptus

- Cymbium with few, less strong spines (fig. 32).............................................................. 2

2(1) Embolic apophysis hooked (fig. 190)........................................................ triangularis sp. n.

2 Embolic apophysis sigmoid or nearly straight ( Figs 4 , 14 , 35 …)................................................. 3

3(2) Embolic apophysis small, occupying only one third of tegulum ( Figs 122 , 136 ); tibial apophysis slender, with narrow base ( Figs 120 , 137 )............................................................................... rhodanicus

- Embolic apophysis large, occupying at least half of tegulum ( Figs 4 , 14 , 35 …); tibial apophysis voluminous, with broad base ( Figs 5 , 12 , 33 …)..................................................................................... 4

4(3) Embolic apophysis with dorsal keel ( Figs 158 , 169 )..................................................... signifer

- Embolic apophysis without dorsal keel ( Figs 4 , 14 , 35 …)...................................................... 5

5(4) Embolic apophysis with 2 prolateral teeth ( Figs 14 , 26 , 93 , 110 )................................................. 6

- Embolic apophysis without ( Figs 4 , 35 , 103 …) or at most with 1 prolateral tooth ( Figs 62 , 129 )....................... 7

6(5) Embolic apophysis slender, elongated and with pointed tip ( Figs 93 , 108 ).................................... omissus

- Embolic apophysis compact, robust, with blunt tip ( Figs 14 , 25 )........................................ dalmatensis

7(6) Embolic apophysis with one prolateral tooth ( Figs 62 , 77 , 129 , 140 )............................................. 8

- Embolic apophysis without prolateral tooth ( Figs 4 , 35 , 103 …)................................................. 9

8(7) Teeth on embolic apophysis equal, apophysis appearing bifid ( Figs 62 , 79 )............................... lyndae sp. n.

- Teeth on embolic apophysis not equal (fig. 129, 142).................................................... rufipes

9(8) Embolus with a strong retrolateral tooth at its base ( Figs 49, 51 )....................................... dentifer sp. n.

- Embolus without retrolateral tooth at base ( Figs 4 , 45 , 103 …)............................................ …….10

10(9) Tibial apophysis nearly symmetric ( Figs 100 , 114 ).................................................. ovatus sp. n.

- Tibial apophysis strongly asymmetric ( Figs 5 , 43 , 147 , 186 )................................................... 11

11(10) Tibial apophysis terminally strongly pointed ( Figs 186 , 195 ).............................................. typhon

- Tibial apophysis bluntly pointed ( Figs 5 , 43 , 147 )........................................................... 12

12(11) Tibial apophysis gradually narrowing, with subterminal concavity ( Figs 43 , 55 )....................... longivulva sp. n.

- Tibial apophysis not gradually narrowing, with basal concavity ( Figs 5 , 147 )..................................... 13

13(12) Embolic apophysis terminally with pointed nipple ( Figs 6 , 20 )........................................... crassipes

- Embolic apophysis terminally blunt ( Figs 149 , 164 )............................................... securifer sp. n. FEMALES

1 Tarsus and tibia of palp with more than 20 ventral spines (fig. 73)..................................... nigroscriptus

- Tarsus of palp with less than 10 ventral spines (fig. 37........................................................ 2

2(1) Fovea triangular ( Figs 178 , 192 )............................................................ triangularis sp. n.

- Fovea oval or rounded ( Figs 7 , 16 , 38 …)................................................................... 3

3(2) Epigyne at least twice as long as wide ( Figs 47 , 64 , 124 )....................................................... 4

- Epigyne less elongate, at most 1.7 times as long as wide ( Figs 7 , 16 , 38 ).......................................... 6

4(3) Fovea narrow anteriorly, widening distinctly to the back ( Figs 64 , 81 )................................... lyndae sp. n.

- Fovea more or less rectangular ( Figs 47 , 124 )............................................................... 5

5(4) Lateral margin of fovea with median angularity ( Figs 47 , 56 )...................................... longivulva sp. n.

- Lateral margin of fovea with parallel margins ( Figs 124 , 138 )........................................... rhodanicus

6(5) Fovea with postero-median septum ( Figs 16 , 95 )............................................................. 7

- Fovea without postero-median septum ( Figs 7 , 38 , 105 )....................................................... 8

7(6) Fovea distinctly longer than wide ( Figs 17 , 28 )..................................................... dalmatensis

- Fovea as long as wide ( Figs 95 , 111 )................................................................. omissus

8(7) Foveal margin with distinct median tooth in lateral margin, dividing fovea into a trapezoid or rectangular anterior part and a rounded posterior part ( Figs 40 , 132 )...................................................................... 9

- Foveal margin without such tooth, fovea oval or rounded ( Figs 7 , 105 , 151 , 160 , 187 ).............................. 10

9(8) Areola anteriorly truncate ( Figs 132 , 143 )............................................................. rufipes

- Areola anteriorly gradually narrowing ( Figs 40 , 52 )................................................ dentifer sp. n.

10(9) Fovea longer than wide ( Figs 105 , 187 )................................................................... 11

- Fovea as long as wide ( Figs 7 , 151 , 160 ).................................................................. 12

11(10) Fovea oval, anterior margins converging ( Figs 105 , 116 ).............................................. ovatus sp. n.

- Fovea not oval, anterior margins diverging ( Figs 187 , 196 )................................................ typhon

12(11) Areola elongated, anteriorly continuing into a chitinised crest ( Figs 160 , 170 )................................. signifer

- Areola less elongated, anteriorly rounded ( Figs 7 , 151 )....................................................... 13

13(12) Epigyne as long as wide ( Figs 151 , 165 ), lateral pockets nearly rounded ( Figs 152 , 166 )................... securifer sp. n.

- Epigyne longer than wide, ( Figs 7 , 22 ), lateral pockets more oval ( Figs 9 , 23 )............................... crassipes

Figures

FIGURES 1–9. Haplodrassus crassipes (Algeria, 1,3‒6: male from Mansourah, 2: female from Djébel Babor; 7. female from Djébel Bouthaleb; 8‒9: female from Djébel Ouarsenis). 1. Male, dorsal aspect; 2. Female, dorsal aspect; 3. Male left palp, retrolateral view; 4. Idem, ventral view; 5. Male palpal tibia, dorsolateral view; 6. Embolic apophysis of male palp, ventral view; 7. Epigyne, ventral view, specimen without mating plugs; 8. Idem, specimen with mating plugs; 9. Vulva, ventral view.

FIGURES 10–17. Haplodrassus dalmatensis (France, male from Saint-André, female from Île de Bagaud). 10. Male, dorsal aspect; 11. Female, dorsal aspect; 12. Male palpal tibia, dorsolateral view; 13. Male left palp, retrolateral view; 14. Idem, ventral view; 15. Idem, prolateral view; 16. Epigyne, ventral view; 17. Vulva, ventral view. Male palp (Fig. 14): E=Embolus; MA=Median apophysis; RTA: Retrolateral tibial apophysis; EA=Embolic apophysis. Epigyne (Figs 16, 17): A=Areola; F=Fovea; H=Hood; LP=Lateral pocket; PF=Pro-fovea; S=Septum; Sp=Spermatheca.

FIGURES 30–40. Haplodrassus dentifer sp. n. (paratypes from Morocco, Aoulouz). 30. Male, dorsal aspect; 31. Female, dorsal aspect; 32. Cymbium of male palp, dorsal view; 33. Male palpal tibia, dorsolateral view; 34. Male left palp, retrolateral view; 35. Idem, ventral view; 36. Idem, retrolateral view; 37. Distal part of female palp, lateral view; 38‒39. Epigyne, ventral view, two different specimens; 40. Vulva, ventral view.

FIGURES 118–125. Haplodrassus rhodanicus (male from Portugal, Alcochete, female from France, Leucate). 118. Male, dorsal aspect; 119. Female, dorsal aspect; 120. Male palpal tibia, retrolateral view; 121. Male left palp, retrolateral view; 122. Idem, ventral view; 123. Idem, prolateral view; 124. Epigyne, ventral view; 125. Vulva, dorsal view.

FIGURES 136–144. Figs 136‒139. Haplodrassus rhodanicus (male from Portugal, Alcochete, female from France, Leucate). 136. Male left palp, ventral view; 137. Male palpal tibia, dorsolateral view; 138. Epigyne, ventral view; 139. Vulva, ventral view. Figs. 140‒144. Haplodrassus rufipes (Lucas) (male from France, Saint-André, female from Spain, Torremolinos). 140. Male left palp, ventral view; 141. Male palpal tibia, dorsolateral view; 142. Embolic apophysis, ventral view; 143. Epigyne, ventral view; 144. Vulva, ventral view.

FIGURES 154–161. Haplodrassus signifer (male from Crete, Nida Plateau, female from France, Corse). 154. Male, dorsal aspect; 155. Female, dorsal aspect; 156. Male left palp, retrolateral view; 157. Idem, ventral view; 158. Embolic apophysis, ventral view; 159. Male palpal tibia, dorsolateral view; 160. Epigyne, ventral view; 161. Vulva, dorsal view.

FIGURES 162–171. Figs 162‒166. Haplodrassus securifer sp. n. (Tunisia, Gafsa). 162. Male left palp, ventral view; 163. Male palpal tibia, dorsolateral view; 164. Embolic apophysis, ventral view; 165. Epigyne, ventral view; 166. Vulva, ventral view. Figs 167‒171. Haplodrassus signifer (male from Crete, Nida Plateau, female from France, Corse). 167. Male left palp, ventral view; 168. Male palpal tibia, dorsolateral view; 169. Embolic apophysis, ventral view; 170. Epigyne, ventral view; 171. Vulva, ventral view.

FIGURES 18–29. Figs 18‒23. Haplodrassus crassipes (Algeria, 18‒21: male from Mansourah, 22‒23: female from Djébel Bouthaleb). 18. Male left palp, retrolateral view; 19. Idem, ventral view; 20. Embolic apophysis of male palp, ventral view; 21. Male palpal tibia, dorsolateral view; 22. Epigyne, ventral view; 23. Vulva, ventral view. Figs 24‒29. Haplodrassus dalmatensis (Algeria, Chréa). 24. Male left palp, retrolateral view; 25. Idem, ventral view; 26. Embolic apophysis of male palp, ventral view; 27. Male palpal tibia, dorsolateral view; 28. Epigyne, ventral view; 29. Vulva, ventral view.

FIGURES 87–97. Haplodrassus omissus (Algeria, male from Chréa, female from Aïn Oghrab). 87. Male, dorsal aspect; 88. Female, dorsal aspect; 89. Female, ventral aspect; 90. Embolic apophysis of male palp, ventral view; 91. Male palpal tibia, dorsolateral view; 92. Male left palp, retrolateral view; 93. Idem, ventral view; 94. Idem, prolateral view; 95. Epigyne, ventral view; 96. Vulva, ventral view; 97. Vulva, dorsal view.

FIGURES 108–117. Figs. 108‒112. Haplodrassus omissus (Algeria, male from Chréa, female from Aïn Oghrab). 108. Male left palp, ventral view; 109. Male palpal tibia, dorsolateral view; 110. Embolic apophysis, ventral view; 111. Epigyne, ventral view; 112. Vulva, ventral view. Figs 113‒117. Haplodrassus ovatus sp. n. (male from Algeria, Biskra, female from Tunisia, Nefzana). 113. Male left palp, ventral view; 114. Male palpal tibia, dorsolateral view; 115. Embolic apophysis, ventral view; 116. Epigyne, ventral view; 117. Vulva, ventral view.

FIGURES 98–107. Haplodrassus ovatus sp. n. (male from Algeria, Biskra, female from Tunisia, Nefzana). 98. Male, dorsal aspect; 99. Female, dorsal aspect; 100. Male palpal tibia, dorsolateral view; 101. Embolic apophysis of male palp, ventral view; 102. Male left palp, retrolateral view; 103. Idem, ventral view; 104. Idem, prolateral view; 105. Epigyne, ventral view; 106. Vulva, ventral view; 107. Vulva, dorsal view.

FIGURES 58–66. Haplodrassus lyndae sp. n. (Algeria, Col Telmet). 58. Male, dorsal aspect; 59. Female, dorsal aspect; 60. Male palpal tibia, dorsolateral view; 61. Male left palp, retrolateral view; 62. Idem, ventral view; 63. Idem, prolateral view; 64. Epigyne, ventral view; 65. Vulva, ventral view; 66. Vulva, dorsal view.

FIGURES 126–135. Haplodrassus rufipes (male from France, Saint-André, females from Spain, Torremolinos). 126. Male, dorsal aspect; 127. Female, dorsal aspect; 128. Male left palp, retrolateral view; 129. Idem, ventral view, embolus partly broken off; 130. Male palpal tibia, dorsolateral view; 131. Embolic apophysis of male palp, ventral view, embolus partly broken off; 132. Epigyne, ventral view, specimen without concretions; 133. Vulva, dorsal view; 134. Epigyne, specimen with concretions; 135. Idem, vulva, ventral view.

FIGURES 77–86. Figs 77‒82. Haplodrassus lyndae sp. n. (Algeria, Col Telmet). 77. Male left palp, ventral view; 78. Male palpal tibia, dorsolateral view; 79. Embolic apophysis of male palp, ventral view; 80. Embolus, dorsal view; 81. Epigyne, ventral view; 82. Vulva, ventral view. Figs 83‒86. Haplodrassus nigroscriptus (type series from Morocco, Essaouira). 83. Male left palp, ventral view; 84. Male palpal tibia, dorsolateral view; 85. Epigyne, ventral view; 86. Vulva, ventral view.

FIGURES 49–57. Figs 49‒53. Haplodrassus dentifer sp. n. (paratypes from Morocco, Aoulouz). 49. Male left palp, ventral view; 50. Male palpal tibia, dorsolateral view; 51. Embolic apophysis of male palp and base of embolus, ventral view; 52. Epigyne, ventral view; 53. Vulva, ventral view. Figs 54‒57. Haplodrassus longivulva sp. n. (Morocco, Missour). 54. Male left palp, ventral view; 55. Male palpal tibia, dorsolateral view; 56. Epigyne, ventral view; 57. Vulva, ventral view.

FIGURES 41–48. Haplodrassus longivulva sp. n. (Morocco, Missour). 41. Male, dorsal aspect; 42. Female, dorsal aspect; 43. Male palpal tibia, dorsolateral view; 44. Male left palp, retrolateral view; 45. Idem, ventral view; 46. Idem, prolateral view; 47. Epigyne, ventral view; 48. Vulva, dorsal view.

FIGURES 145–153. Haplodrassus securifer sp. n. (Tunisia, Gafsa). 145. Male, dorsal aspect; 146. Female, dorsal aspect; 147. Male palpal tibia, dorsolateral view; 148. Male left palp, retrolateral view; 149. Idem, ventral view; 150. Idem, prolateral view; 151. Epigyne, ventral view; 152. Vulva, ventral view. 153. Vulva, dorsal view.

FIGURES 181–189. Haplodrassus typhon (male from France, Bouches-du-Rhône, female from Spain, Granada). 181. Male, dorsal aspect; 182. Female, ventral aspect; 183. Male left palp, retrolateral view; 184. Idem, ventral view; 185. Idem, prolateral view; 186. Male palpal tibia, dorsolateral view; 187. Epigyne, ventral view 188. Vulva, ventral view; 189. Idem, dorsal view.

FIGURES 190–197. Figs. 190‒193. Haplodrassus triangularis sp. n. (Morocco, Sidi Harazem).190. Male left palp, ventral view; 191. Male palpal tibia, dorsolateral view; 192. Epigyne, ventral view; 193. Vulva, ventral view. Figs 194‒197. Haplodrassus typhon (male from France, Bouches-du-Rhône, female from Spain, Granada). 194. Male left palp, ventral view; 195. Male palpal tibia, dorsolateral view; 196. Epigyne, ventral view; 197. Vulva, ventral view.

FIGURES 172–180. Haplodrassus triangularis, sp. n. (Morocco, Sidi Harazem). 172. Female, dorsal aspect; 173. Idem, ventral aspect; 174. Male left palp, retrolateral view; 175. Idem, ventral view; 176. Idem, prolateral view; 177. Male palpal tibia, dorsolateral view 178. Epigyne, ventral view 179. Vulva, ventral view; 180. Idem, dorsal view.