Domiporta austrogallica ( Mayer-Eymar, 1898 ) Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard, 2021, The Mitridae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4983 (3), pp. 1-72 : 30

Domiporta austrogallica ( Mayer-Eymar, 1898) nov. comb.

Figs 8B 1 –B 2

Mitra scrobiculata Brocc. — Hörnes 1852b: 100, pl. 10, fig. 18 [non Brocchi, 1814].

* [ Mitra suballigata ] Varietät austro-gallica M.-E.— Mayer-Eymar 1898: 82 [nov nom pro. Hörnes, 1852b, pl. 10, fig. 18].

Mitra (Tiara) orientalis var. latisulcata n. var. —Kojumdgieva in Kojumdgieva & Strachimirov 1960 (pars): 156 [non pl. 41, figs 12–14 = Domiporta amoena nov. sp.].

Type material. Holotype: NHMW 1846 /0037/0098, SL: 31.4 mm, MD: 8.0 mm, Baden ( Austria), illustrated in Hörnes (1852b, pl. 10, fig. 18), Figs 8B 1 –B 2 .

Revised description. Shell medium sized, very slender fusiform with deeply incised suture. Protoconch and first teleoconch whorl unknown. Teleoconch of about eight whorls. Spire whorls weakly convex, nearly straight sided, with periphery along mid-whorl. Early teleoconch whorl cancellate, with four convex spiral cords. Spiral cords forming two pairs, separated by distinct central spiral groove with rectangular cross-section on subsequent whorls. Spiral grooves between pairs of spiral cords successively becoming obsolete abapically, resulting in two broad spiral bands. Third spiral band hardly visible along abapical suture. Last whorl with ten spiral bands, abapically narrowing, separated by prominent spiral grooves. Aperture elongate, narrow. Columellar callus indistinct. Columella with moderately prominent, oblique adapical columellar fold, weak middle fold and subobsolete abapical one. Siphonal canal long, relatively wide, straight with deeply incised siphonal notch.

Shell measurements and ratios. SL = 31.4 mm, MD: 8.0 mm, AA = 25°, SL/ MD: 4.01, AL/AW: 7.4, AH/S: 1.9.

Discussion. Mayer-Eymar (1898) introduced Mitra suballigata var. austrogallica for the Austrian specimen, illustrated by Hörnes (1852b, pl. 10, fig. 18). Kojumdgieva in Kojumdgieva & Strachimirov (1960) was obviously not aware of the paper by Mayer-Eymar (1898) and proposed Mitra orientalis latisulcata as name for the same specimen, which is thus an objective junior synonym. The specimens illustrated by Kojumdgieva in Kojumdgieva & Strachimirov (1960), however, are not conspecific with Domiporta austrogallica , but represent Domiporta amoena nov. sp.

Cancilla suballigata (Bellardi, 1887) was described from the Tortonian of Stazzano ( Italy) and differs from Domiporta austrogallica in its much wider last whorl, narrower spiral grooves and it lacks the amalgamating spiral cords on the spire whorls (see syntype in Bellardi 1887b, pl. 4, fig. 15; Ferrero-Mortara et al. 1981, pl. 48, figs 12a–b and Serravallian specimens in Landau et al. 2013, pl. 33, figs 12–13). Domiporta amoena nov. sp. is less slender, has more spiral cords, its periphery is more convex and the mode of spiral cord formation is splitting instead of amalgamating. Domiporta paucicostata (Bellardi, 1887) , from the Langhian of the Monte dei Cappuccini (Colli Torinesi, Italy), seems to represent a closely related species (see Bellardi 1887a: 70, pl. 4, fig. 12). It is distinguished from D. austrogallica by its broader shell, presence of four spiral cords on the penultimate whorl and the higher number of spiral bands on the last whorl.

Palaeoenvironment. The clay of the Baden Formation formed in middle to outer neritic settings with up to 250 m water depth ( Hohenegger et al. 2008).

Distribution in Central Paratethys. Badenian (middle Miocene): Vienna Basin: Baden ( Austria). Note that Kojumdgieva in Kojumdgieva & Strachimirov (1960) referred to Viennese specimens and that her illustrated specimens represent Domiporta amoena nov. sp.

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Palaeogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser & Landau 2017) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements for shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 8. A1–A2. Domiporta turpis nov. sp., holotype, Baden (Austria), NHMW 1846/0037/0098a. B1–B2. Domiporta austrogallica (Mayer-Eymar, 1898), holotype, Baden (Austria), NHMW 1846/0037/0098. C1–C2. Domiporta amoena nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0106/0002. D1–D2. Domiporta amoena nov. sp., holotype, Lăpugiu de Sus (Romania), NHMW 2020/0106/0001. E1–E2. Domiporta amoena nov. sp., paratype, Lăpugiu de Sus (Romania). F1–F2. Domiporta amoena nov. sp., paratype, Lăpugiu de Sus (Romania). G1–G2. Domiporta pulchra nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0129/0003. H1–H2. Domiporta pulchra nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0129/0004. I1–I2. Domiporta pulchra nov. sp., holotype, Lăpugiu de Sus (Romania), NHMW 2020/0129/0001.