Dissochaeta vacillans (Blume) Blume, Flora 14: 495. 1831.

Kartonegoro, Abdulrokhman, Veldkamp, Jan Frits, Hovenkamp, Peter & Welzen, Peter van, 2018, A revision of Dissochaeta (Melastomataceae, Dissochaeteae), PhytoKeys 107, pp. 1-178 : 1

53. Dissochaeta vacillans (Blume) Blume, Flora 14: 495. 1831. Fig. 27 , Map 24

Melastoma vacillans Blume, Bijdr. Fl. Ned. Ind. 17: 1074. 1826.

Dissochaeta fusca Blume, Flora 14: 497. 1831. Type: Indonesia. Java, C.L. Blume 1791 (lectotype, designated by Kartonegoro and Veldkamp 2010, pg. 143: L [L0729468]!; isolectotypes: K [K000859621]!, L [L0537244]!, P [P05283572, image seen]!,).

Dissochaeta fusca Blume var. ferruginea Blume, Flora 14: 497. 1831. Type: Indonesia. Java, Bantam, J.C. van Hasselt s.n. (lectotype, designated by Kartonegoro and Veldkamp 2010, pg. 143: L [L0537248]!; isolectotypes: L [L0537239, L0537247]!).

Dissochaeta fusca Blume var. obtuso-acuminata Blume, Flora 14: 497. 1831. Type: Indonesia. West Java: Buitenzorg, Tjiampea, C.L. Blume s.n. (lectotype, designated by Kartonegoro and Veldkamp 2010, pg. 143: L [L0537242]!; isolectotypes: L [L0537246]!).

Dissochaeta velutina Blume, Flora 14: 497. 1831. Type: Indonesia. Java, Bantam, Leuwi Boengoer, H. Kuhl & J.C. van Hasselt s.n. (lectotype, designated by Kartonegoro and Veldkamp 2010, pg. 143: L [L0537234]!; isolectotypes: K [K000859622]!, L [L0537249, L0537250]!).

Dissochaeta decipiens Blume, Mus. Bot. 1(3): 36. 1849. Type: Indonesia. Java, H. Kuhl & J.C. van Hasselt s.n. (lectotype, designated by Kartonegoro and Veldkmap 2010, pg. 131: L [L0008892]!; isolectotypes: K [K000859489]!, L [L0008891]!).

Dissochaeta inappendiculata Blume var. fusca (Blume) Miq., Fl. Ned. Ind. 1(1): 525. 1855.

Dissochaeta monticola auct. non Blume: Triana, Trans. Linn. Soc. London 28: 83. 1872. p.p., excl. type.

Neodissochaeta fusca (Blume) Bakh. f., Contr. Melastom.: 136. 1943.

Neodissochaeta vacillans (Blume) Bakh. f., Contr. Melastom.: 144. 1943.

Type.

Indonesia. West Java: Buitenzorg, Tjiawi, C.G.C. Reinwardt s.n. (lectotype, designated by Kartonegoro and Veldkamp 2010, pg. 143: L [L0008894]!; isolectotype: L [L0008895]!).

Description.

Climbing up to 3 m height. Branchlets terete or subquadrangular, 2-6 mm diameter, glabrescent to stellate-furfuraceous; nodes swollen, with interpetiolar ridge; internodes 4.5-10 cm long. Leaves: petioles terete, 5-24 mm long, puberulous to furfuraceous; blades ovate, ovate-oblong or elliptic-oblong, 6.5-15 × 2.5-7 cm, membranous, base rounded, margin entire, apex acuminate, tip 0.5-2 cm long; nervation with 1 pair of lateral nerves and 1 pair of intramarginal nerves; adaxially glabrous, abaxially glabrous to nearly brown sparsely stellate-furfuraceous. Inflorescences terminal and in the upper leaf axils, up to 30 cm long, many-flowered; main axis angular, sparsely stellate-furfuraceous; peduncle up to 7 cm long; primary axes up to 25 cm long with 4-6 nodes, secondary axes up to 4 cm long with 1-3 nodes, tertiary axes up to 1.5 cm long with 1 node; bracts minute or linear, ca. 3 mm long, furfuraceous, caducous; bracteoles minute or linear to oblong, 1.5-4 mm long, stellate-furfuraceous; pedicels sparsely stellate-furfuraceous, 4-6 mm long in central flowers, 1-3 in lateral flowers. Hypanthium campanulate or suburceolate, 2-5 × 1-3 mm, glabrescent or sparsely to densely stellate-furfuraceous; calyx lobes truncate, 0.5-1 mm long, with 4 undulating apices, rounded, or subtriangular, glabrous; petal buds conical, 2-4 mm long; mature petals ovate or oblong, 5-6 × ca. 3 mm, base clawed, apex obtuse, glabrous or with minute hairs at margin, white-pinkish or pink. Stamens 8, sometimes 4 with the oppositipetalous ones undeveloped, unequal when 8, filaments straight; alternipetalous stamens with 2-4 mm long filaments, anthers oblong or lanceolate, thecae 3-5 mm long, yellow, pedoconnective ca. 0.5 mm long, basal crests triangular, 1-2 mm long, margin irregular, lateral appendages paired, filiform, 1.5-3 mm long; oppositipetalous stamens when developed with 2-3 mm long filaments, anthers ovate-oblong or lanceolate, thecae 2.5-4 mm long, basal crests triangular or ligular, 0.5-1 mm long, erect, lateral appendages paired or reduced to a single lateral one, filiform, 1-2 mm long. Ovary half or ⅔ of hypanthium in length, apex puberulous or pubescent; style 5-8 mm long, apex curved, glabrous; stigma minute; extra-ovarial chambers 8, the 4 alternipetalous ones extending between the apex and the middle of the ovary, the 4 oppositipetalous ones shallow. Fruits subglobose to urceolate, 3 –6(– 10) × 2-6 mm, glabrous; stalks 2 –5(– 7) mm long, calyx lobe persistent, erect. Seeds ca. 0.5 mm long.

Distribution.

Java and Lesser Sunda Islands (Sumbawa).

Ecology and habitat.

Forest, secondary or depleted forest or edge of river at 500-1400 m elevation.

Vernacular names.

Java: harendong areuy, harendong bokor areuy, harendong gede (Sunda).

Note.

Dissochaeta decipiens , with only four fertile, alternipetalous stamens, is synonymised with D. vacillans , because it has a similar appearance due to the indumentum on the branchlets, leaves and inflorescences; moreover, the shape of the stamens and the appendages are also similar.

Specimens examined.

INDONESIA. Banten: Pandeglang, Mt. Karang, C.A. Backer 7470 (BO); Ibid., Galusur, 500 m, 28 May 1912, S.H. Koorders 40659β (BO); Ibid., 700 m, 1 Jun 1912, S.H. Koorders 40727β (BO); Ibid., Menes, Mt. Pulasari, 1000 m, Mar 1913, C.A. Backer 7055 (BO); Lebak, J.C. van Hasselt s.n. (L); Ibid., Leuwi Bungur, H. Kuhl & J.C. van Hasselt s.n. (L); Ibid., Citorek & Muncang, C.A. Backer 1835 (BO). Central Java: Mt. Slamet, Baturaden, 1000 m, 30 Mar 1970, B.J. Bernardius D43051 (BO); Pekalongan, Josorejo, C.A. Backer 16219 (BO). West Java: Bogor, Mt. Salak, Gunung Bunder to Kawah Ratu, 1300 m, 8 Jan 1941, C.N.A. de Voogd & S. Bloembergen s.n. (BO); Ibid., C.A. Backer 4198 (BO); Ibid., Upper Lido, 1200 m, 22 Feb 2000, H. Wiriadinata & W.S. Hoover 31188 (BO); Ibid., W.S. Hoover & M. Hendra 32564 (BO); Leuwiliang, Puraseda, 500 m, 2 Feb 1929, C.G.G.J. van Steenis 2712 (BO); Ibid., Mt. Butik Buligir, C.A. Backer 6150 (BO); Ibid., Mt. Sunarari, 1000 m, 1 Jan 1913, C.A. Backer 6380 (BO); Ciampea, C.L. Blume s.n.(L); Ciawi, C.G.C. Reinwardt s.n. (L); Puncak Pass, Tugu, Above Gunung Mas, 1300-1500 m, 18 Mar 1952, W. Meijer 105 (BO, K, L); Ibid., Gunung Luhur, Tugu, 1700 m, 8 Aug 1982, M.M.J. van Balgooy & J.P. Mogea 4284 (BO); Mount Pangrango, Bodogol, A. Kartonegoro 318 (BO); Mt. Halimun, C.A. Backer 10914 (BO); Ibid., R.C. Bakhuizen van den Brink 3336 (BO, L, U); Ibid., Cikaniki, 1000 m, 11 Jan 2001, D. Arifiani et al. 142 (BO); Ibid., 9 Mar 2000, W.S. Hoover, D. Girmansyah & J. Hunter 32172 (BO); Ibid., Nirmala, 1300 m, 10 Jun 1980, M.M.J. van Balgooy & H. Wiriadinata 2922 (BO, L); Ibid., Malasari, 1055 m, 10 Oct 2017, A. Kartonegoro 1105 (BO); Cianjur, Mt. Gede, Cibodas, 1450 m, J.G. Boerlage s.n. (BO); Ibid., R.H.C.C. Scheffer s.n. (BO); Ibid., 3 Nov 1987, E.A. Widjaja 3220 (BO); Ibid., Sindanglaya, 1000 m, Dec 1916, C.A. Backer 21507 (BO); Ibid., Pasir Pangsalatan, 1500 m, 2 Jun 1948, Enoh 181 (BO, L); Ibid., S.H. Koorders 31670β (BO); Sukanegara, E.R. Hellendoorn 5 (BO); Cibeber, Campaka, 1000 m, 16 Jun 1923, J.J. Smith 822 (BO, L); Ibid., Cidadap, Cadas Malang, 1000 m, 19 Apr 1916, R.C. Bakhuizen van den Brink 1469 (BO); Ibid., 20 Mar 1923, W.F. Winckel 1176β (BO,K, L); Takokak, 1200 m, 9 Jun 1900, S.H. Koorders 33314β (BO); Ibid., S.H. Koorders 33315β (BO); Bandung, Cigenteng, 1400 m, 26 Jan 1897, S.H. Koorders 26306β (BO); Mt. Tangkuban Prahu, 1600 m, 26 Jul 1927, W.M. Docters van Leeuwen 11487a (BO); Mt. Sembung, Margalangu, 1200 m, 19 Mar 1914, C.A. Backer 12298 (BO); Garut, Rawa Cangkuang, R.H.C.C. Scheffer s.n. (BO); Ibid., Pasawahan, 400 m, 31 Dec 1911, C.A. Backer 2261 (BO); Ibid., Mt. Ciparay, 1100 m, 27 Jul 1914, C.A. Backer 15041 (BO); Mt.Cikuray, Pasir Kolotok, 1000 m, 15 Aug 1913, C.A. Backer 8685 (BO); Tasikmalaya, Panjalu, 720 m, 4 Aug 1917, S.H. Koorders 47851β (BO). West Nusa Tenggara. Sumbawa, Sumbawa Barat, Mt. Batulante, 700 m, 3 Nov 1961, A.J.G.H. Kostermans 19164 (BO, K, P).

Figures

Map 24. Distribution of D. pubescens (●), D. rectandra (■) and D. vacillans (▲).

Figure 27. Dissochaeta vacillans. a habit b branchlet c hypanthium d flowers e immature fruit. Photographs by A. Kartonegoro; voucher: Kartonegoro 1105 (BO).