Dissochaeta beccariana Cogn. in A.DC. & C.DC., Monogr. Phan. 7: 559. 1891.

Kartonegoro, Abdulrokhman, Veldkamp, Jan Frits, Hovenkamp, Peter & Welzen, Peter van, 2018, A revision of Dissochaeta (Melastomataceae, Dissochaeteae), PhytoKeys 107, pp. 1-178 : 1

9. Dissochaeta beccariana Cogn. in A.DC. & C.DC., Monogr. Phan. 7: 559. 1891. Map 8

Neodissochaeta magnibracteata Bakh. f., Contr. Melastom.: 142. 1943. Type: Indonesia. West Kalimantan: Soengei Kenepai, 1893, J.G. Hallier 2013 (holotype: L [L0537263]!; isotypes: BO [BO1865993, BO1865994, BO1865995]!, K [K000859505]!, SING n.v.).

Neodissochaeta beccariana (Cogn.) M.P.Nayar, Kew Bull. 20: 159. 1966.

Type.

Malaysia. Sarawak: Santubong, O. Beccari PB 2190 (lectotype, designated here: FI [FI007928, image seen]!; isolectotype: K [K000859504]!).

Description.

Climbing up to 30 m in height. Branchlets terete, 3-5 mm in diameter, glabrous, sometimes with scattered bristles; nodes swollen, with a raised interpetiolar, crest-like ridge, sparsely covered with stellate hairs; internodes 8-9 cm long. Leaves: petioles terete, 1.3-1.5 cm long, glabrous except small bristles abaxiallyly; blades broadly ovate, 11-16 × 6.2-7.5 cm, subcoriaceous, base rounded, margin entire, apex acuminate, tip up to 2 cm long; nervation with 1 pair of lateral nerves and 1 pair of intramarginal nerves; adaxially glabrous, abaxially glabrous with sparsely stellate hairs on the midrib, with a basal pair of glandular patches. Inflorescences terminal, up to 22 cm long, many-flowered; main axis glabrous to sparsely stellate-puberulous; primary axes up to 18 cm long with 4 or 5 nodes, secondary axes up to 4 cm long with 1 or 2 nodes, tertiary axes ca. 1 cm with 1 node; bracts elliptic, ca. 3 × 1.2 cm, conspicuous, glabrous, white; bracteoles ovate, conspicuous, 6-8 × 6-7 mm, glabrous, white; pedicels glabrous or sparsely stellate-puberulous, 5-6 mm long in central flowers, 2-3 mm long in lateral flowers. Hypanthium tubular, 4-7 × 2-3 mm, glabrous or stellately-puberulous; calyx lobes truncate, with or without 4 small undulate tips, ca. 0.5 mm long; petal bud conical, up to 4 mm long, apex with narrow acute tip; mature petals broadly ovate, 6-7 × 5-6 mm, reflexed, purple, base clawed to slightly cordate, apex acute. Stamens 8, subequal, filaments straight, light yellow; alternipetalous stamens with 4-5 mm long filaments, anthers curved, sickle-shaped, thecae 4-5 mm long, apex acute, purple, pedoconnective ca. 1 mm long, basal crest triangular, thin, erose, irregular margin 1-2 mm long, lateral appendages paired, filiform, 3-5 mm long; oppositipetalous stamens with 5-6 mm long filaments, bent at the attachment to anthers, anthers thick, slightly curved, hook-shaped, thecae 5-7 mm long, yellow, apex obtuse, basal crest thin, ligular, 1-2 m long, lateral appendages paired, filiform, 1-2 mm long. Ovary half as long as hypanthium, apex glabrous; style glabrous, 16-18 mm long, curved at the end, slender, pale yellow; stigma minute, pinkish; extra-ovarial chambers 8, shallow to nearly undeveloped. Fruits subglobose to ovoid, 5-8 × 3-6 mm, glabrous, with 8 slight lines along the surface; calyx lobe remnants persistent. Seeds ca. 0.75 mm long.

Distribution.

Borneo.

Ecology and habitat.

Lowland to montane dipterocarp forest in open shade at 30-1400 m elevation.

Vernacular name.

Akar (Iban).

Note.

The species is easily recognised because all parts are usually glabrous, the leaf blades abaxiallyly have a pair of glandular patches at the base and the bracts and bracteoles are white. The most similar species is D. glandulosa , also with glabrous parts and glandular patches, but it has a larger, campanulate hypanthium.

Selected specimens examined.

MALAYSIA. Sabah: Beaufort, Beaufort Rail-Line, 10 m, 22 Mar 1962, G. Mikil SAN 34540 (K, L); Keningau, Pensiangan Kayu, 17 Oct 1985, Sumbing SAN 110362 (L); Ibid., Crocker Range, 1400 m, 17 Oct 1999, S.J. Davies et al. 99128 (L); Tawau, Road Kalabakan, 26 Jul 1962, Aban Gibot SAN 30555 (K, L); Ranau, Mt. Kinabalu, Dallas, 900 m, 5 Aug 1931, J. Clemens & M.S. Clemens 30342 (BM, K, L). Sarawak: O. Beccari PB 2190 (FI, K); Sibu, Rejang, Jul 1893, G.D. Haviland 3628 (BM, K); Bario, Bukit Lawi, 1500 m, 2 Aug 1985, D. Awa & B. Lee S.50537 (L); Kuching, Gunung Penrissen, 2 May 1962, Ilias Paie S.16326 (K); Ibid., Semenggoh FR., 19 Oct 1966, Banyeng ak Nyudong S.26207 (BO, K, L); Belaga, Linau Balui, Sg. Nawai, 800 m, 5 Sep 1978, B. Lee S.40000 (K, L); Kapit, Balleh, Menyiong, Sungai Sebatu, 500 m, 12 Nov 1979, Othman bin Haron et al. S.41368 (K, L); Ibid., Wong Kijang, 25 Oct 1988, Othman et al. S.56049 (L); Limbang, Sungai Ensungei, Tg. Long Amok, 11 Sep 1980, R. George et al. S.42845 (K, L); Miri, Gunung Mulu National Park, 1400 m, 9 Mar 1990, P.C. Yii & Abu Talib S.58277 (K, L). BRUNEI. Belait: Melilas, Ulu Ingei, 30 m, 5 Mar 1996, Said BRUN 17315 (K). Temburong: Ulu Belalong, 500 m, 21 Jan 1994, J. Dransfield et al. 7392 (K, L). INDONESIA. East Kalimantan: West Kutai, Long Ibut, 130 m, 16 Aug 1925, F.H. Endert 2552 (BO, L); Ibid., Mount Kemul, 1200 m, 26 Sep 1925, F.H. Endert 3572 (BO, L). West Kalimantan: Pontianak, Gunung Bentuang, 200 m, 24 Jun 1989, J.S. Burley & Tukirin 2857 (BO, L); Sungai Kenepai, 1893, J.G. Hallier 2013 (BO, K, L); Sintang, 120 m, 18 Apr 1994, A.C. Church et al. 1000 (BO, L); Kapuas Hulu, Mendalam Rivers, 170 m, 15 Mar 2000, Albertus 45 (L).

Figures

Map 8. Distribution of D. beccariana (●), D. brassii (■) and D. celebica var. celebica (▲).