Diplommatina calcarata Möllendorff, 1897

Hausdorf, Bernhard, 2017, Revision of the Diplommatinidae (Gastropoda: Cyclophoroidea) from Java, Zootaxa 4312 (2), pp. 201-245 : 220-221

Diplommatina calcarata Möllendorff, 1897

( Figs 14 , 31 ; Table 1)

Diplommatina calcarata Möllendorff, 1897: 94 (JaVa) . Van Benthem Jutting 1948: 596, fig. 53. Heryanto 2009: 332. Heryanto 2011: 47, figS.

Diplommatina (Sinica) calcarata — Kobelt 1902: 458. Zilch 1953: 27, pl. 13 fig. 189.

Diagnosis. Diplommatina calcarata is characterized by the distinct narrowing of the last 1.5 whorls, distinct extensions of the peristome, a prominent columellar lamella, widely spaced ribs and a strongly ascending last whorl.

Shell ( Fig. 31 ; Table 1). Dextral; spindle-shaped with conical early whorls and a much narrowed last whorl; with 5.5–6.75 whorls separated by a deep suture; protoconch smooth; teleoconch with widely spaced ribs (15–28 on the penultimate whorl; approximately the same number on the last whorl); without discernible spiral striae between the ribs; whitish; subtranslucent; last whorl rounded, with a distinct lateral right constriction; aperture subangular, angular at the transition from columellar to basal side; upper insertion of the peristome distinctly ascending towards aperture; insertions of the peristome connected by a detached callus; peristome distinctly expanded, with 2–4 blunt processes, of which the upper palatal and the columellar-basal are strongest, thickened, doubled; aperture with prominent columellar lamella, forming a high lamella inside the last whorl; with a long parietal lamella laterally left to laterally right inside the last whorl; inside the last whorl ventrally with a short spiral palatal fold above the middle of the whorl and with a radial palatal fold laterally right; umbilicus closed; operculum thin, corneous, circular, paucispiral. Shell height 2.10–2.65 mm, shell diameter 1.3–1.7 mm.

Remarks. Diplommatina calcarata differs from D. tetragonostoma in the smaller, more widely ribbed shell and the last whorl that ascends strongly towards the aperture.

Habitat. In leaf litter, rocks and deadwood in forests on calcareous and volcanic soils at 80–1700 m a.s.l.

Type material. Indonesia, Java (lectotype of D. calcarata [designated by Zilch 1953] SMF 105207; paralectotypes of D. calcarata SMF 105208/2).

Other material. Indonesia, Java, Banten: Lebak, Sawarna, Mt. Kembang, 81 m a.s.l., 6°58'21"S 106°16'46"E (MZB 18793/8; ZMH 133002/6).—West Java: Bogor, Dungus Iwul, 100 m a.s.l., 6°31'S 106°24'E (ZMA 317631/ 6); Bogor, Leuwiliang, near waterfall of the Cianten, 300 m a.s.l., 6°38''S 106°38'E (ZMA 68406/8, vouchers to van Benthem Jutting 1948); Bogor, Ciampea, Mt. Cibodas, 6°33'S 106°41'E (MZB 5919/10); Sukamantri, Mt. Salak, 6°41'S 106°45'E (BDJ/1); Bogor Botanical Garden, 6°35'S 106°47'E (ZMA 317626/1); Depok, 150 m a.s.l., 6°24'S 106°49'E (ZMA 68405/5, vouchers to van Benthem Jutting 1948); Bogor, Puncak Pass, northern edge of Telaga Warna, 1500 m a.s.l., 6°42'S 107°00'E (MZB 4058/3; ZMA 68404/1, vouchers to van Benthem Jutting 1948); Cianjur, Mt. Gede, 1200 m a.s.l., 6°44'S 107°00'E (ZMA 33714/3); Cianjur, Pancoran Mas, near Rarahan, Mt. Gede, 1400 m a.s.l., 6°43'S 107°00'E (ZMA 68403/7, vouchers to van Benthem Jutting 1948); Cianjur, Cibodas, along path behind Pesanggrahan, 1500 m a.s.l., 6°45'S 107°00'E (SMF 105209/2; ZMA 317623/2, 317625/4); Cianjur, Cibodas, Botanical Garden, 1400 m a.s.l., 6°45'S 107°00'E (ZMA 317618, 317619/1, 317621/ 3, 317622/1, 317624/1, 317627/1, 317628/1, 317630/6, 317652/1); Cibodas, 1400 m a.s.l., 6°45'S 107°01'E (MZB 4059/1; ZMA 317620/4, 317629/1; van Benthem Jutting 1948); Cianjur, Cibeureum, 1400 m a.s.l., 6°45'S 107°01'E (MZB 4057/1); Sukabumi, Cibadak, Pasir Salam, 1224 m a.s.l., 7°02'S 107°03'E (BDJ/1); Mt. Tilu W of Pangalengan, 7°10'S 107°34'E (JV 3842/3); Bandung, Mt. Malabar, 1600 m a.s.l., 7°07'S 107°36'E (ZMA 68407/ 1); Garut, Talaga Bodas, 1700 m a.s.l., 7°12'36''S 108°04'01''E (MZB 18011/4); Palutungan, 2000–2150 m a.s.l., 6°55'S 108°24'50"E (Heryanto 2009); Palutungan, 1850–2000 m a.s.l., 6°55'20"S 108°24'40"E (Heryanto 2009); Palutungan, 1700–1850 m a.s.l., 6°55'20"S 108°25'E (Heryanto 2009); Palutungan, 1550–1700 m a.s.l., 6°55'40"S 108°25'E (Heryanto 2009); Palutungan, 1400–1550 m a.s.l., 6°56'S 108°25'20"E (Heryanto 2009); Linggajati, 1550–1700 m a.s.l., 6°53'20"S 108°26'10"E (Heryanto 2009); Linggajati, 1400–1550 m a.s.l., 6°53'20"S 108°26'20"E (Heryanto 2009); Linggajati, 1250–1400 m a.s.l., 6°53'20"S 108°26'30"E (Heryanto 2009); Linggajati, 1100–1250 m a.s.l., 6°53'20"S 108°26'40"E (Heryanto 2009); Kuningan, Mt. Ciremai, 1500 m a.s.l., 6°53'S 108°26'E (ZMA 68402/1, vouchers to van Benthem Jutting 1948).—East Java: Bremi near Mt. Argopuro, 1200 m a.s.l., 7°58'S 113°29'E (JV 4015/8).

Distribution ( Fig. 14 ). Diplommatina calcarata is probably endemic to Java. A re-examination of specimens recorded as D. calcarata from Borneo ( Vermeulen 1993) proved that they belong to a separate species. A record from Sumatra (Heryanto 2013) also needs to be checked.

Figures

FIGURES 10 – 14. DiStribution of Diplommatinidae in JaVa. 10. Diplommatina planicollis Möllendorff, 1897, Diplommatina heteroglypha Van Benthem Jutting, 1948 and Diplommatina javana Möllendorff, 1897. 11. Diplommatina halimunensis n. sp., Diplommatina kakenca n. sp. and Diplommatina heryantoi n. sp. 12. Diplommatina perpusilla Möllendorff, 1897 and Diplommatina ristiae n. sp. 13. Diplommatina sulcicollis Möllendorff, 1897 and Diplommatina cyclostoma Möllendorff, 1897. 14. Diplommatina calcarata Möllendorff, 1897 and Diplommatina tetragonostoma Möllendorff, 1897.

FIGURES 29 – 32. Diplommatina Spp. from JaVa. 29. Diplommatina perpusilla Möllendorff, 1897, JaVa (lectotype SMF 105210). 30. Diplommatina ristiae n. sp., Central JaVa: Donorejo Village, near Seplawan CaVe, KaligeSing, Purworejo (holotype MZB 18886). 31. Diplommatina calcarata Möllendorff, 1897, JaVa (lectotype SMF 105207). 32. Diplommatina tetragonostoma Möllendorff, 1897, WeSt JaVa: Bandung, Mt. Wayang (holotype SMF 105199). Scale bar = 1.0 mm.