Cymbasoma javense, Grygier & Suárez-Morales, 2018

Grygier, Mark J. & Suárez-Morales, Eduardo, 2018, Recognition and partial solution of nomenclatural issues involving copepods of the family Monstrillidae (Crustacea: Copepoda: Monstrilloida), Zootaxa 4486 (4), pp. 497-509 : 505-506

Cymbasoma javense sp. nov.

Holotype (data from Suárez-Morales 2000:144–145): undissected adult male in ethanol, Museum für Naturkunde, Berlin ( Germany), sample no. 26279, Bangka Strait, western Java Sea, Indonesia, labelled as “ Leg Prinz Adalbert, Dr. Sanders. Det. M.J. Isaac, 1973”.

Description: See Suárez-Morales (2000: 145–146, figs. 11–26).

Diagnostic remarks [for a more detailed treatment, see Suárez-Morales (2000: 148–149)]: Uniquely among the Monstrillidae , a dorsal hump is present anteriorly on the cephalothorax; four pairs of rounded processes (the first pair bare, the members of the other three pairs each bearing a row of subrectangular scale-like processes) are lined up in two longitudinal rows between the bases of the antennules; and antennular setal elements 3 and IVv have peculiar projecting sockets that are ornamented respectively with many short spinules and many small, ovoid plates. Also unusual are the absence of setules on the natatory setae of legs 1–4 and on the inner margin of the terminal spiniform seta of exopodal segment 3 in these legs. The male genitalia differ from those of Monstrilla wandelii Stephensen, 1913 sensu Park (1967) , with which Isaac (1974a) noted resemblances, in having lappets with simple, not bifid, distal ends and in the absence of fifth legs.

Cymbasoma lenticula Suárez-Morales & McKinnon, 2016 . The specific name is spelled two ways in its original description by Suárez-Morales & McKinnon (2016), mostly as C. lenticula , but also as C. lentilum on p. 95. We take this opportunity to act as First Revisers under Article 24.2.3 of the Code and fix C. lenticula as the correct original spelling.

Monstrilla capitellicola Hartman, 1961 . Suárez-Morales et al. (2010) did not recognize M. capitellicola as valid and regarded it as being of uncertain taxonomic status because Hartman (1961) based it on immature females still enclosed in a capsular sheath. Validity is a concept that is only relevant in the context of competing names. Rather than “invalid”, it would be more accurate to regard M. capitellicola as a nomen dubium, or as a nomen inquirendum if the possibility exists of clarifying its identity through examination of type or newly collected material, including by rearing to adults.

Monstrilla mariaeugeniae Suárez-Morales & Islas-Landeros, 1993 . Suárez-Morales (1998) had two nomenclaturally valid ways to handle the supposed synonymy of male Monstrilla wandelii tropica Suárez- Morales, 1996 and female M. mariaeugeniae Suárez-Morales & Islas-Landeros, 1993 from the Mexican Caribbean. He chose to retain M. mariaeugeniae as a full species with M. w. tropica as its junior subjective synonym. His other option was to reduce the senior taxon to the subspecies M. wandelii mariaeugeniae (again with M. w. tropica as its junior subjective synonym) along with the nominotypical subspecies M. w. wandelii Stephensen, 1913 , known from Greenland and the Canadian and Alaskan Pacific. The question of recognizing subspecies among the relevant females was not addressed, however, and this latter option was not mentioned. Suárez-Morales (2001a) later synonymized male M. w. tropica with female M. elongata Suárez-Morales, 1994 instead of M. mariaeugeniae , so the point is now mostly moot, but the taxonomic (and eventual nomenclatural) question of the status of M. mariaeugeniae vis à vis M. wandelii , i.e. as a separate species or a subspecies, remains.

Monstrillopsis boonwurrungorum Suárez-Morales & McKinnon, 2014 . The specific name is spelled two ways in its original description by Suárez-Morales & McKinnon (2014), mostly as M. boonwurrungorum , but also as M. boonwurrungi in the second to last line of the Remarks on p. 311. We take this opportunity to act as First Revisers under Article 24.2.3 of the Code and fix M. boonwurrungorum as the correct original spelling.

Monstrillopsis reticulata ( Davis, 1949) . Grygier & Ohtsuka (2008) advised that resolution of the problem posed by assignment of the specific name reticulata to supposedly non-conspecific males and females [see Suárez- Morales et al. (2006) for details] requires designation of a lectotype (p. 501). In fact, the latter authors had already examined the syntypes of M. reticulata but could not make any use of them because the specimens were nearly invisible on the slides. A request to the International Commission on Zoological Nomenclature to replace the present syntypes by a neotype (of either sex, preferably a new specimen from the type locality, Biscayne Bay, Florida), to fix the applicability of the name to a single taxon, therefore seems to be required.