Ctenodontina baleta ( Walker, 1849 ) Walker, 1849

Vieira, Rodrigo, Landa, José Manuel Ayala & Rafael, José Albertino, 2017, Ctenodontina Enderlein, 1914 (Diptera, Asilidae, Asilinae): new combinations, synonyms, new species and new records for Argentina, Zootaxa 4216 (3), pp. 279-291 : 285-289

publication ID	https://doi.org/ 10.5281/zenodo.231832
publication LSID	lsid:zoobank.org:pub:6E036E6B-0D91-482A-AAB3-DDA147837FA3
DOI	https://doi.org/10.5281/zenodo.6010910
persistent identifier	https://treatment.plazi.org/id/211B878C-FFEB-FF84-FF28-E62F0229FDEA
treatment provided by	Plazi
scientific name	Ctenodontina baleta ( Walker, 1849 )
status	comb. nov.

Treatment

Ctenodontina baleta ( Walker, 1849) comb. nov.

( Figs. 6 View FIGURES 2 – 9 , 24–26 View FIGURES 24 – 28. 24 , 34–47 View FIGURES 34 – 41 View FIGURES 42 – 47 )

Asilus baletus Walker, 1849: 427 View in CoL ; Schiner, 1866: 721; Williston, 1891: 90 (catalogue); Kertész, 1909: 323 (catalogue); Hull, 1962 (2): 545; Martin & Papavero, 1970: 88 (catalogue).

Lecania baleta ; Artigas & Papavero, 1995: 37; Papavero, 2009: 31 (catalogue).

Pachychaeta ? caracasae Martin, 1975: 85 View in CoL ; Artigas & Papavero, 1995: 37; Papavero, 2009: 31 (catalogue).

Pachychoeta caracasae Martin, 1975: 85 View in CoL syn. nov. ( Figs. 27 View FIGURES 24 – 28. 24 –29)

Pachychoeta inca Martin, 1975: 86 View in CoL syn. nov. ( Figs. 30–33 View FIGURES 30 – 33 )

Addenda to the holotype original description based on specimens from Venezuela.

Male terminalia ( Figs. 38–44 View FIGURES 34 – 41 View FIGURES 42 – 47 ): Black. Medioapical region of hypandrium with concavity ( Fig. 39 View FIGURES 34 – 41 ). Epandrium wide basally with finger-like lobe placed dorsally. Apex of finger-like lobe rounded in lateral view ( Fig. 40 View FIGURES 34 – 41 ); subepandrial sclerite without lateral projections, apical margin rounded ( Fig. 41 View FIGURES 34 – 41 ); gonocoxite with apical 1/3 digitiform ( Fig. 42 View FIGURES 42 – 47 ); gonostyle subrectangular, covered with small setae anteriorly and with long setae posteriorly ( Figs. 42, 43 View FIGURES 42 – 47 ); aedeagus with two long prongs ( Fig. 44 View FIGURES 42 – 47 ); ejaculatory apodeme wide in lateral view ( Fig. 44 View FIGURES 42 – 47 ). Female terminalia ( Fig. 46, 47 View FIGURES 42 – 47 ): Tergite VIII expanded laterally ( Figs. 46, 47 View FIGURES 42 – 47 ); tergite IX+X slightly membranous medially ( Fig. 46 View FIGURES 42 – 47 ); apex of sternite VIII truncate with mediaoapical slit ( Fig. 47 View FIGURES 42 – 47 ); hypoproct with spiniform setae; median sclerite at posterior end of furca (remnant of tergite IX) absent; two oval sclerotized spermathecal capsules.

Comments. Ctenodontina baleta (Walker) comb. nov., Pachychoeta caracasae Martin syn. nov. and Pachychoeta inca Martin syn. nov. were allocated in Lecania Macquart. However , these species have diagnostic characters of Ctenodontina . Lecania is a paraphyletic group and has no diagnostic characters defined. According to Artigas & Papavero (1995), Lecania “is an extremely varied assembly of species. This group needs a thorough revision to decide whether there are several genera involved or only one. The variation of the several characters is amazing, and there is an incredible combination of them”.

Ctenodontina baleta (Walker) comb. nov., P. caracasae Martin View in CoL syn. nov., P. inca Martin View in CoL syn. nov. as well as C. nairae View in CoL and C. martini View in CoL , do not have a spinous swelling ventrally on male hind femora ( Figs. 6, 7 View FIGURES 2 – 9 ). However, as discussed by Fisher (1985) and Vieira (2012b), Ctenodontina View in CoL is better defined by its very distinctive male and female terminalia and C. baleta (Walker) comb. nov., P. caracasae Martin View in CoL syn. nov. and P. inca Martin View in CoL syn. nov. by the following characters: female ovipositor with an apical comb of spines, and tergite VIII broadly expanded laterally, densely setose and cupped beneath, completely obscuring sternite VIII in profile and male specimens with an epandrium narrowed apically (usually terminating in a finger-like lobe), distinctly shorter than sternite VIII ( Fisher & Hespenheide 1992; Fisher 2009; Vieira 2012b).

Material examined. Holotype of Asilus baletus : Venezuela / One of Walkers series so named. Eaw / Type Pachychaeta baleta (Wlk) [Walker] , N. Papavero det. / Holotype Asilus baletus Walker , det. J.E. Chainey, 1985 (♂ NHM); Holotype of Pachychaeta caracasae : Venezuela, 5.of Caracas, Quebrada Catia 18.xi.1968 / Chas. H. Martin Collector / California Academy of Sciences, Type No. 12602 / Pachychaeta caracasae Martin, Det. C.H. Martin / Holotype Pachychaeta caracasae Chas. H. Martin (♂ CAS) ; Holotype of Pachychaeta inca: San Esteban , Venezuela, 1–15.Dec.1939 / P. Anduze Collector / California Academy of Sciences, Type No. 12603 / Holotype Pachychaeta inca Chas. H. Martin / Pachychoeta inca Martin, Det. C.H. Martin 19 (♂ CAS) .

Other material examined. [ Venezuela], Distrito Federal, Rio Oritapo , 200m, col.: M. Ayala- 11.6.[19]67 (1 ♂, 1 ♀ INPA) ; [ VENEZUELA], 7-x-1967, Edo. Guárico, La Puerta, [San Francisco de Tiznados-Qda. Coronado 09°31’05"N 67°37’29"W], col. Adolfo Ayala (2 ♂, 2 ♀ INPA) GoogleMaps ; [ Venezuela], rio Oritapo , Edo. Vargas: (10.621446°, -66.480845°) / 22-VIII- [19]65 (2 ♂ JMAC) ; same data, except: 26-XI- [19]65 (1 ♂ JMAC) ; same data, except: 11- VI- [19]67 (2 ♂, 3 ♀ JMAC) ; [ Venezuela] rio Camurí Grande , Edo. Vargas: (10.613400°, -66.719940°) / 23-IX- [19]72 (1 ♀ JMAC) ; [ Venezuela] rio Cojo Macuto- Edo. Vargas: (10.599056°, -66.881881°) / 9-VIII- [19]69 (1 ♂ JMAC) ; [ Venezuela] rio Curupao , Edo. Miranda: (10.497547°, -66.644678°) / 10-VI- [19]67 (1 ♂, 1 ♀ JMAC) ; [ Venezuela] rio Suapire, El Manguito , Edo. Miranda: (10.294581°, -66.737645°) / 6-I- [19]73 (1 ♀ JMAC) ; [ Venezuela] rio Periquito, La trilla, Edo. Aragua: (10.388835°, -67.745727°) / 15-XII- [19]73 (1 ♂, 1 ♀ JMAC) ; same data, except: 12-X- [19]68 (3 ♂, 2 ♀ JMAC) ; [ Venezuela] rio Maletero, La Victoria , Edo. Aragua: (10.236862°, -67.315002°) / 2-IX- [19]77 (3 ♀ JMAC) ; [ Venezuela] rio Guárico - San Sebastian - La Puerta, Edo . Aragua: (9.916317°, -67.349211°) / 13-VII- [19]67 (1 ♀ JMAC) ; same data, except: 7-X- [19]67 (10 ♂, 6 ♀ JMAC) ; [ Venezuela] rio Aragua, La Victoria, Pié de Cerro : (10.256486°, -67.348120°) / 18-V- [19]74 (1 ♂, ♀ JMAC) ; same data, except: 29-VI- [19]74 (1 ♂, ♀ JMAC) ; [ Venezuela] Hacienda Carrizalito - Palambrota , Edo. Guárico: (9.651826°, -67.552502°) / 20-XII- [19]65 (1 ♀ JMAC) ; [ Venezuela] rio Orituco , Edo. Guárico: (9.910627°, -66.394536°) / 11-XI- [19]72 (1 ♂, 1 ♀ JMAC) ; [ Venezuela] rio Borburata , Edo. Carabobo: (10.425188°, -67.977241°) / 9-VIII- [19]86 (1 ♂, 1 ♀ JMAC) .