Mniotype bathensis (Lutzau, 1900)

Volynkin, Anton V., Matov, Alexei Yu., Behounek, Gottfried & Han, Hui-Lin, 2014, A review of the Palaearctic Mniotype adusta (Esper, 1790) species-group with description of a new species and six new subspecies (Lepidoptera: Noctuidae), Zootaxa 3796 (1), pp. 1-32 : 12

Mniotype bathensis (Lutzau, 1900)

( Figs 35–38 , 75, 76 , 92 , 112 , 113 )

Crino bathensis Lutzau, 1900 , Entomologische Zeitschrift, Frankfurt am Main 14: 162 (Type locality: [ Germany]).

Synonymy: Hadena adusta var. moesta Staudinger, 1897 (Type locality: [ Russia, Transbaikalia, Yablonovoi Ridge] «Apfelgebirge») syn. n.; Crino adusta subsp. urupino Bryk, 1942 (Type locality: Russia, Kurile islands, «Urup, Tokotambai») syn. rev. Polia urupolia Bryk, 1942 (Type locality: Russia, Kurile islands, «Urup, Tokotanbai»); Blepharita hoenei Sugi, 1959 (Type locality: Japan, Nokkio Yumoto), nec. Heydemann, 1938.

Type material examined. Lectotype ( Figs 36 , 112 ). Designated herein for Hadena adusta var. moesta Staudinger, 1897 : female, " adusta var. moesta Stgr. " / "Apfelgeb., Sibir. or., 96 Dörr." / pink label "Origin" / "58" / "ex coll. Staudinger 112" / " MNHU " / (the lectotype deposited in coll. NKMB). Slide GB7870f Behounek.

Other matherial examined: 84 males, 123 females from Finland, NE Germany, Switzerland, Estonia, Latvia, Russia (Karelia, Saint-Petersburg, Novgorod, Tver, Jaroslavl, Moscow, Vladimir, Perm, Ekaterinburg, Orenburg, Tomsk, Omsk, Altai, Kemerovo, Krasnojarsk, Irkutsk, Chita and Kamchatka areas, Tatarstan, Buryatia, Khakassia, Altai Republic, Kurile Islands), North Mongolia (Töv and Bulgan aimaks) (Colls ZISP, SZMN, AVB, ASV, BBT).

Slides AV0513, AV0515, AV0686, AV0687, AV0688, AV0689, AV0706, AV0707, AV0711, AV0712, AV0734, AV0735, AV0765, AV1124 Volynkin (males), AV0514, AV0636, AV0637, AV0638, AV0639, AV0699, AV0700, AV0703 Volynkin, OP2278f Pekarsky (females).

Notes. The taxon moesta has been treated by some authors (Hacker 1990; Ronkay et al. 2001) as a Siberian subspecies of M. adusta . Examination of the genitalia of the lectotype of Hadena adusta var. moesta ( Fig. 112 ) showed that it is conspecific with M. bathensis . The taxon moesta is a monotonous brownish grey form of M. bathensis occuring in East Palaearctic. The taxon urupino was treated by Ronkay et al. (2001) as a subspecies of M. bathensis , but Kononenko (2005) treated it as a junior synonym of bathensis . By the examination of the material from Kurile Islands and Kamchatka, it was found that the reddish form described by Bryk (1942) as urupino ( Figs 37, 38 ) occurs together with darker form common in Europe ( Fig. 35 ) and the monotonous brownish grey form described as moesta ( Fig. 36 ). Thus, Pacific populations of M. bathensis cannot be treated as a distinct subspecies.

Diagnosis. Wingspan 35–49 mm. The external differences of M. bathensis are described in diagnoses of M. adusta and M. kobyakovi . The male genitalia ( Figs 75, 76 , 92 ) differ from M. adusta in the somewhat narrower juxta with shorter serrated parts of its apical processes (in M. bathensis distal third, but in M. adusta distal half serrate), larger elongated cucullus with longer corona, more curved digitus, apically obtuse carina, shorter eversible dorso-lateral bar beginning from apex of carina and having longer teeth basally; and from M. kobyakovi in the somewhat narrower juxta with shorter serrated parts of its apical processes (in M. bathensis distal third, but in M. kobyakovi distal half serrate), larger elongated cucullus with longer corona, more curved digitus, shorter eversible dorso-lateral bar of carina with longer teeth basally. The female genitalia ( Figs 112 , 113 ) differ from M. adusta in the narrower ostium bursae with a concave posterior margin, narrower ductus bursae and smaller signum; and from M. kobyakovi in the narrower ostium bursae with a concave posterior margin, and somewhat narrower corpus bursae.

Distribution. Eurasiatic Palaearctic. North Europe, NE European part of Russia, Ural, South Siberia, Transbaikalia, Mongolia, Russian Far East (including Kamchatka and Kurile islands), Northeastern China (Manchuria), Korea, Japan.

Figures

FIGURES 31 – 40. Mniotype spp. adults: 31, M. kobyakovi, holotype male, Russia, Altai Mts (ZISP); 32, ditto, paratype male, Russia, Altai Mts (AVB); 33, ditto, paratype male, S Siberia, Tyva (MDS) (Photo by M. Dvořák); 34, ditto, paratype male, China, Inner Mongolia (ASV) (Photo by A. Saldaitis); 35, M. bathensis, male, Russia, Leningrad territory (ZISP); 36, ditto, female, lectotype of Hadena adusta var. moesta, East Siberia, Yablonovoi Ridge (NKMB); 37, ditto, female, Kurile Islands (ZISP); 38, ditto, male, same locality (ZISP); 39, M. aulombardi, paratype female, Taiwan (GBG / ZSM); 40, ditto, male, Taiwan (HMHN).

FIGURES 75 – 80. Mniotype spp., male genitalia: 75, M. bathensis, Russia, Altai Mts, slide AV 0513 Volynkin; 76, M. bathensis, Russian Far East, Kurile Islands, slide AV 0735 Volynkin; 77, M. aulombardi, Taiwan, slide 0 259 Matov; 78, M. aulombardi, Taiwan, slide AV 0786 Volynkin; 79, M. anilis anilis, France, Alpes-Maritimes, slide AV 0789 Volynkin; 80, M. a. sylvatica, France, Corsica, slide AV 0788 Volynkin.

FIGURES 87 – 96. Mniotype spp., male genitalia: carina. 87, M. lama lama, Kyrgyzstan, slide AV 0681 Volynkin; 88, M. adusta adusta, E Kazakhstan, slide AV 0713 Volynkin; 89, M. adjuncta adjuncta, Pakistan, slide AV 0792 Volynkin; 90, M. juldussica, Tajikistan, slide AV 0796 Volynkin; 91, M. kobyakovi, paratype, Altai Mts, slide AV 0744 Volynkin; 92, M. bathensis, Russia, Altai Mts, slide AV 0513 Volynkin; 93, M. aulombardi, Taiwan, slide AV 0786 Volynkin; 94, M. anilis anilis, Italy, Reg. Lazio, slide GB 8141 m Behounek; 95, M. dubiosa dubiosa, E Kazakhstan, slide AV 0680 Volynkin; 96, M. krisztina, China, Tibet, slide hhl- 2506 - 1 Han.

FIGURES 105 – 112. Mniotype spp., female genitalia: 105, M. adjuncta adjuncta, Himalaya Mts, slide AV 0776 Volynkin; 106, M. a. cineritia, paratype, Nepal, slide AV 0791 Volynkin; 107, M. juldussica, neotype, Central Tien Shan Mts, slide 0 328 Matov; 108, ditto, lectotype of Mamestra vicina, slide 0 276 Matov; 109, ditto, Tajikistan, Gissarsky Ridge, slide AV 0697 Volynkin; 110, M. kobyakovi, paratype, Altai Mts, AV 0722 Volynkin; 111, ditto, paratype, Altai Mts, slide AV 0766 Volynkin; 112, M. bathensis, lectotype of Hadena adusta var. moesta, slide GB 7870 f Behounek.

FIGURES 113 – 120. Mniotype spp., female genitalia: 113, M. bathensis, South Siberia, slide AV 0725 Volynkin; 114, M. aulombardi, paratype, Taiwan, slide 8144 f Behounek; 115, M. anilis anilis, France, slide GB 8142 f Behounek; 116, M. a. sylvatica, Corsica, slide AV 0787 Volynkin; 117, M. dubiosa dubiosa, paralectotype, " Mustagata Yarkend ", slide 7659; 118, ditto, SE Kazakhstan, slide AV 0878 Volynkin; 119, M. d. amitayus, paratype, W China, Tibet Plateau, slide AV 0777 Volynkin; 120, M. krisztina, China, Tibet, slide hhl- 2271 - 2 Han.