Cretolyra shawi, Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot, 2024

Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo & Perrichot, Vincent, 2024, Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities, Arthropod Systematics & amp; Phylogeny 82, pp. 151-181 : 151

3.1.1.1.2. † Cretolyra shawi sp. nov.

Figures 2B , 4

Etymology.

The specific epithet is a patronym honoring Scott R. Shaw, for his contributions to the knowledge of Megalyridae .

Material studied.

Holotype female NIGP203545; housed in the Nanjing Institute of Geology and Paleontology (NIGP), Chinese Academy of Sciences, Nanjing, China.

Type locality.

Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar.

Age.

Upper Albian to lower Cenomanian, mid-Cretaceous.

Diagnosis.

Pedicel bilobed, long and thick (Figs 4D-E ; vs. calciform and elongate in Cretolyra noijebumensis gen. et sp. nov.); axillae almost rounded and reduced on edges of mesoscutellum, not contiguous medially (vs. contiguous medially in Cretolyra noijebumensis gen. et sp. nov.); axillar grooves smooth (Fig. 4A ; vs. axillar grooves crenulated in Cretolyra noijebumensis gen. et sp. nov.); medial cell nearly triangular, with 1m-cu very short (Fig. 2B ; vs. medial cell rectangular, with 1m-cu distinct in Cretolyra noijebumensis gen. et sp. nov.).

Description.

Body length 2.82 mm. - Head globular, higher than long (length ca. 0.40 mm), covered with short setae; frons convex, smooth, divided by sulcus; compound eye oval, higher than long; vertex convex; clypeus smooth, apically rounded; mandibles symmetrical, with three teeth; toruli separated from each other by less than their own diameter; subantennal groove present; antenna about half body length; scape shorter than pedicel; pedicel thick, bilobed; flagellomeres cylindrical, longer than wide; flagellomere 1 shorter than flagellomeres 2-5 (length 0.15 mm vs. ca. 0.21 mm); flagellomeres 6-11 shorter (length ca. 0.10 mm); flagellomere 12 as long as flagellomere 1 (length 0.15 mm); occipital carina minutely crenulate. - Mesosoma one third of body length (length 0.98 mm); mesoscutum convex, divided by smooth median mesoscutal sulcus; parapsidal lines present; axillae small, almost rounded and not contiguous medially; axillar groove smooth; mesoscutellum smooth, convex, diamond-shaped; pronotum smooth, not visible dorsally, with posteromedial part moderately high as viewed laterally; anterior thoracic spiracle not surrounded by pronotal cuticle posteriorly; mesometapectal sulcus crenulate; propodeum carinate. - Fore wing hyaline and covered with microtrichiae, two thirds of body length (length 2.57 mm); pterostigma reduced; C, Sc+R, M+Cu, A, Rs fully pigmented; R1 extending beyond marginal cell; Rs closing marginal cell in straight line; M pigmented to apex; medial cell small, elongate and rectangular, nearly triangular by side of 1m-cu. - Legs with two mesotibial and one metatibial spurs; metacoxa elongate; metafemur and metatibia swollen; metabasitarsus two to three times as long as remaining metatarsomeres; row of comb-like setae along ventral surface of metatibia and metabasitarsus. - Metasoma longer than mesosoma (length 1.44 mm); elongate and narrowed at apex; tergites smooth; hypopygium well-developed; ovipositor shorter than metasoma (length 1.33 mm; OL/BL ratio 0.43), sheaths fully preserved, transversely striated.

Figures

Figure 2. Drawings of fore wing venation for the taxa described in the present paper. A † Cretolyra noijebumensis gen. et sp. nov.; B † Cretolyra shawi gen. et sp. nov.; C † Genkyhag innebula gen. et sp. nov.; D † Megacoxa chandrahrasa gen. et sp. nov.; E † Megacoxa janzeni gen. et sp. nov.; F † Megacoxa synchrotron gen. et sp. nov.; G † Kamyristi exfrigore gen. et sp. nov.; H † Kamyristi yantardakhensis gen. et sp. nov. Scale bars: 0.5 mm (A - H).

Figure 4. † Cretolyra shawi gen. et sp. nov., holotype female NIGP 203545, mid-Cretaceous, Kachin amber. A habitus in dorsal view; B habitus in latero-ventral view; C head and mesosoma in lateral view; D head in frontal view; E line drawing of left scape, pedicel and flagellomeres 1 and 2. Scale bars: 1 mm (A, B); 0.5 mm (C, D, E). Abbreviations: F 1 = flagellomere 1; F 2 = flagellomere 2; MeS = mesometapectal sulcus; pe = pedicel; sc = scape; sp = anterior thoracic spiracle.