Cretolyra noijebumensis, Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot, 2024

Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo & Perrichot, Vincent, 2024, Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities, Arthropod Systematics & amp; Phylogeny 82, pp. 151-181 : 151

3.1.1.1.1. † Cretolyra noijebumensis sp. nov.

Figures 2A , 3

Etymology.

The specific epithet is an adjective referring to the type locality where the amber pieces were collected.

Material studied.

Holotype male IGR.BU-067; housed in the amber collection of the Geology Department and Museum of the University of Rennes, France (IGR).

Type locality.

Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar.

Age.

Upper Albian to lower Cenomanian, mid-Cretaceous.

Diagnosis.

Antennae inserted on edge of clypeus (Fig. 3A ; vs. inserted above clypeal margin in Cretolyra shawi gen. et sp. nov.); axillae contiguous medially and axillar grooves crenulated (Fig. 3B ; vs axillae separated medially and axillar grooves smooth in Cretolyra shawi gen. et sp. nov.).

Description.

Body length 3.33 mm; body very pubescent, especially on metasoma where setae conceal margins of tergites; the setae have agglomerated small bubbles that make observation difficult. - Head globular, higher than long (length 0.55 mm; height 0.75 mm); frons convex, smooth, divided by thin median sulcus; compound eye oval, higher than long; vertex convex, with smooth ocellar triangle; clypeus well-developed; torulus inserted closer to clypeus than to eye; shallow but distinct subantennal groove adjoining ventral margin of eye; antenna half as long as body, almost reaching metasoma; scape twice as long as wide (length 0.16 mm); pedicel shorter than scape (length 0.11 mm); flagellomeres cylindrical, elongate, longer than wide; flagellomere 1 shortest (length 0.15 mm); following flagellomeres longer (length ca. 0.21 mm); occipital carina crenulate. - Mesosoma almost half body length (length 1.29 mm; height 0.65 mm); mesoscutum convex, shagreened, divided by smooth median mesoscutal sulcus (mesoscutum length 0.58 mm; width ~0.60 mm); parapsidal lines present, diverging anteriorly; axillae contiguous medially, axillar groove crenulate; mesoscutellum diamond-shaped; pronotum smooth, not visible dorsally, with posteromedial part moderately high as viewed laterally; anterior thoracic spiracle not fully surrounded by pronotal cuticle; mesometapectal sulcus crenulate; propodeum shorter than mesoscutellum, with incomplete pairs of median and submedian carinae, two complete lateral longitudinal carinae, anterior region of propodeum with transversal row of foveae adjacent to concealed metanotum, posterior region of propodeum with three large foveae. - Fore wing hyaline and covered with microtrichiae, about two thirds of body length (length 2.50 mm); C, Sc+R, M+Cu, A, Rs fully pigmented; R1 extending beyond marginal cell; Rs closing marginal cell in straight line; M pigmented to apex; medial cell rectangular, narrow; Cu almost reaching posterior wing margin. - Legs covered with setae; two mesotibial and one metatibial spurs; metacoxa elongate; metafemur and metatibia swollen; metabasitarsus three times as long as following metatarsomeres and with row of short comb-like setae along ventral surface. - Metasoma half body length (length 1.48 mm), elongate and narrowed at apex; seven smooth tergites with pubescent posterior margin; first tergite largest; second tergite one quarter of metasoma length; remaining tergites shorter.

Figures

Figure 2. Drawings of fore wing venation for the taxa described in the present paper. A † Cretolyra noijebumensis gen. et sp. nov.; B † Cretolyra shawi gen. et sp. nov.; C † Genkyhag innebula gen. et sp. nov.; D † Megacoxa chandrahrasa gen. et sp. nov.; E † Megacoxa janzeni gen. et sp. nov.; F † Megacoxa synchrotron gen. et sp. nov.; G † Kamyristi exfrigore gen. et sp. nov.; H † Kamyristi yantardakhensis gen. et sp. nov. Scale bars: 0.5 mm (A - H).

Figure 3. † Cretolyra noijebumensis gen. et sp. nov., holotype male IGR. BU- 067, mid-Cretaceous, Kachin amber. A habitus in lateral view; B head and mesosoma in dorsal view; C fore wing; D anterior part of mesosoma in lateral view with structures outlined; E metasoma and posterior part of the mesosoma in lateral view. Scale bars: 0.5 mm (A, B, C, E); 0.25 mm (D). Abbreviations: AG = inner axillar groove; MeS = mesometapectal sulcus; MMS = median mesoscutal sulcus; OC = occipital carina; PpL = parapsidal line; sp = anterior thoracic spiracle.