Craspedophorus bonnyi Bates, 1890

Häckel, Martin, 2016, A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 1. Revision of the Craspedophorus reflexus group (Coleoptera: Carabidae), Zootaxa 4061 (5), pp. 504-526 : 507-508

2. Craspedophorus bonnyi Bates, 1890

(Plate 3, Figs 17, 18)

Craspedophorus bonnyi Bates, 1890: 480 (type loc. “R[iver]. Aruwimi” [=Democratic Congo: Orientale Province: forest region towards Albert Lake]). Alluaud 1915: 154, Burgeon 1930: 159, 1935b: 181. Häckel et Farkač 2012: 79.

Type material. Holotype (♀): “R. / Aruwimi [handwritten, white label] // Craspedophorus / Bonnyi / Bates [handwritten, white label] // Ex-Musaeo / H. W. Bates / 1892 [printed, white label] // TYPE [printed, red label] // Muséum Paris // ex Coll. / R. Oberthür / 1952 [printed, blue label]”, (Plate 3, Fig. 18, MNHN, Oberthür/Bates Collection).

Other material examined. Democratic Congo. 1 ♂: “Urwald Beni [=Nord-Kivu Province]”, (Plate 3, Fig. 17, cMH); Error. 1♀: “S-India, Madras state, Kurumbagaran, ex col. Hlinkel, 2011 [erroneous locality]”, (cIB).

Note. The original description of this species is based on a single specimen collected by William Bonny, who was a military man in Stanley's expedition.

Description (in part, see Bates 1890: 480). “Larger body, head relatively smaller than pronotum, elytra wider and convex. Black, glossy, head convex, weakly constricted behind eyes; pronotum almost ovoid, with almost rectangular posterior angles, base flat, lateral margins posteriorly sinuate and elevated, quite coarsely punctured and sparsely covered by setae; elytra smooth, striated, punctured, with intervals pubescent quite sparsely and finely for this genus, each with two transverse straw-coloured maculae extending from 4th to 8th interval… ventrites anteriorly not crenulated, [met]episterna square, slightly longer than wide” [from Latin]. “Length 23 mm. In the proportions of head and thorax to the pterothorax similar to C. eximius Laferté [= C. reflexus ( Fabricius, 1781) ], with which it also agrees in the moderate strangulation of the neck and in the suboval outline of the thorax, which has, however, more sinuated sides behind the middle, and distinct, almost rectangular, hind angles. The elytra, in the single example which appears not to be abraded, is nearly glabrous, the lateral interstices only having a few hairs; the striae are deep and punctured and the punctuation of the interstices is rather shallow and nowhere very dense: the fasciae are straw-yellow, narrow, and macular, the spot on the fifth interstice of both fasciae being much shorter than the others; on the hinder fascia the spot on the sixth interstice is extended forward and that on the seventh prolonged behind.” The coloration of C. bonnyi most closely resembles C. bozasi Alluaud, 1930 , from which it differs by pronotal shape (more transverse, sharper and more projecting hind angles), elytral sculpture (striae more finely punctate), and the length and width of humeral maculae (Plate 3, cf. Figs 17, 18 and 19). From other species of the group near C. reflexus it differs by shape of pronotal hind angles (sharp, more projecting) and shape of the maculae, namely humeral. The shape of the antennae (not flattened), front margins of ventrites (not crenulate) and the metepisterna place this species in the C. reflexus group, near species with two orange maculae on each elytron (Plates 1, 3). The differential characters are given in the key to species presented below.

Distribution. Democratic Congo: Orientale and Nord- Kivu Provinces.

Bates (1890: 479) noted. “Mr. Bonny informs me that they [specimens] were all taken at Yambuya Camp and on the march through the forest-region towards Albert Nyanza, between the months of October 1887 and November 1888 ...”