Coptocarpus cyllodinus (Fauvel, 1882) Will & Guéorguiev, 2021

Will, Kipling & Gueorguiev, Borislav, 2021, Phylogenetic systematics of the genera of Thryptocerina Jeannel, 1949 and new species from New Caledonia (Coleoptera, Carabidae, Oodini), ZooKeys 1044, pp. 375-425 : 375

publication ID	https://dx.doi.org/10.3897/zookeys.1044.63775
publication LSID	lsid:zoobank.org:pub:D50CC77C-2E69-41FD-A9BD-395B025C43AE
persistent identifier	https://treatment.plazi.org/id/2F6758F0-659B-5A2F-B018-4BB2FD61DCAE
treatment provided by	ZooKeys by Pensoft
scientific name	Coptocarpus cyllodinus (Fauvel, 1882)
status	comb. nov.

Treatment

Coptocarpus cyllodinus (Fauvel, 1882) View in CoL comb. nov. Figs 12 View Figures 9–12 , 19 View Figures 18–21 , 24 View Figures 22–31 , 34 View Figures 32–40 , 47 View Figures 46, 47 , 48 View Figure 48 , 50 View Figures 49–51 , 52 View Figures 52, 53

Hoplolenus cyllodinus Fauvel, 1882: 266.

Type locality.

Nouméa.

Material examined.

Holotype: New Caledonia • ♂, card-mounted specimen, original label with “Nouméa” glued to pink label " Coll. R.I.Sc.N.B. Nouvelle Calédonie rec Deplanche ex coll. Fauvel "// “TYPE” // "Coll.et det. A. Fauvel Hoplolenus cyllodinus n. sp. R.I.Sc. N.B. 17.479" [RBINS].

Other material.

New Caledonia • ♂; Mt. Koghis ; 22°11'S, 166°31'E; track entrance; 500 m; 6.v.2006; berleseate; G.B.Monteith; [QM] GoogleMaps • ♀; Mt. Koghis ; 22°11'S, 166°00'E; 750 m; 29.xi.2000; berleseate, sieved litter, rainforest; G.B. Monteith; [QM] GoogleMaps • ♂; 22°10'28"S, 166°30'48"E, 700 m, 12.iii.2007; K. Will; EMEC1075689; [EMEC] • ♂; same data as for preceding; headlamp search; EMEC1075690; [EMEC] GoogleMaps • ♂; same data as for preceding; under rocks/logs; EMEC1075691; [EMEC] GoogleMaps • ♂; Noumea Mt. Kogni; 12-13.II.1977; leg. Dr. J. BALOGH; [NHMUK] • ♀; “Kuakui” [ Mt. Kouakoue ]; P.D. Montague; B. M. 1946-210; [NHMUK] • ♂; Yahoué; “février”; [RBINS, Fauvel collection, but not a syntype] • ♂; Riviere Bleue Park : 22°09'00"S, 166°41'12"E, 330 m, Houp Géant trail, 13.iii.2007, headlamp search, K. Will; EMEC1075688; [EMEC] GoogleMaps • ♀; same data as for preceding; [EMCE] GoogleMaps • ♂; Mt. Dzumac ; 22°03'S, 166°28'E; Mt. Dzumac Road; 700 m; 1.xii.2000; berleseate, sieved litter, rainforest; G.B. Monteith; [QM] GoogleMaps • ♂; same data as for preceding; [ZSM] GoogleMaps • ♀; Col de Yate ; 22°10'S, 166°55'E; 2.xi.2004; hand collecting, CJ Burwell; [QM] GoogleMaps • ♀; Pic du Pin ; 22°14'S, 166°50'E; 280 m; 25-26.xi.2004; day hand collecting, rainforest; QM party; [QM] GoogleMaps .

Diagnosis.

A very distinctive, large species for the New Caledonian fauna with a deep notch below the eye that may receive the antenna (Fig. 50 View Figures 49–51 ); eye and orbit very prominent, almost shelf-like; male metafemur with large, ventral tooth (Fig. 48 View Figure 48 ), clypeal setae absent, elytral stria 6 and 7 impressed throughout, 5 only impressed in apical 1/2, 1-4 not evident or only shallowly impressed near the apex. Aedeagus with distinct, broad apical blade (Fig. 24 View Figures 22–31 ).

Redescription.

Habitus. Large sized for New Caledonia oodines (BL: 7.35-8.45 mm, BW: 3.60-4.40 mm), with elliptic, moderately convex body (Fig. 19 View Figures 18–21 ). Color and luster. Head dark reddish black to black; clypeus and mandibles reddish brown; palpi, antennae and tarsi yellow to yellow brown; pronotum, elytra and ventral surface of body black; legs (excl. tarsi) deep reddish brown to reddish black. Integument moderately shiny, mostly without spectral iridescence, excluding proepisterna and abdominal ventrites on sides where it is present. Microsculpture and punctation. Dorsal surface of head (excluding mandibles) and pronotum, with isodiametric meshes well impressed and visible; elytra with isodiametric meshes weakly visible; prosternum with scarcely-visible isodiametric sculpticells, ventrites 4-6 medially with scarcely-visible slightly transverse meshes, rest of ventral surface with sculpticells mostly not evident. Dorsal surface lacking macropunctation; head, pronotum and elytra with micropunctation, smaller on head and most of pronotum, larger on basal 1/4 of pronotum and elytra; micropunctation regular on metacoxae, very scattered on last abdominal ventrites, absent on rest of ventral surface, abdominal ventrites 1-3 macropunctate on sides, 2-5 finely coarsely wrinkled on sides. Chaetotaxy. Labrum with six setae, each in its own socket, four lateral setae larger than medial two. Clypeal setae absent. Supraorbital setae absent. Elytron without discal setiferous punctures. Abdominal ventrites 1-5 without ambulatory setae. Head. Approximately 2 1/2 × as wide as pronotum (Suppl. material 2: Table S1). Eye size large, eye prominently protruded, EyW/HW: 1.38-1.48. Labrum anterior margin somewhat concave. Frontoclypeal suture evident, but slightly impressed. Antenna relatively short, with pubescence starting from segment 4. Last labial palpomere swollen, blunt at apex, longer and larger than penultimate one. Mentum tooth with paramedial border distinct. Thorax. Pronotum ~ 1 2/3 wider than long (PW/PL: 1.64-1.85); width at apex 2-2 2/3 × less than at widest point (PW/PA: 1.95-2.38). Disc with midline fine, shallowly impressed, and ended well before apical and basal margins, without apical transverse impression; anterior angles weakly projected, rounded at tips, anterior margin concave, submarginal sulcus laterally present, lacking in medial 3/10. Prosternal process rhomboid, elongate, broadly pointed at apex, bordered at apex, not bordered at sides. Mesosternum anteriorly without medial tubercle. Metepisternum ~ 1 1/2 × wider than long, with lateral margin convex, coadunation with epipleuron long, located medially and posteriorly; suture between metepisternum and metepimeron barely distinct. Elytra. Broad, slightly longer than wide (EL/EW: 1.05-1.17). Basal margin forming small, sharp tooth at shoulder, medially nearing parascutellar puncture to which it is joined by impressed basal section of stria 2. Humeral submarginal carina present. Apical sinuation shallow, barely distinct. Elytral striae 6 and 7 entirely impressed, 5 impressed only at posterior 1/3, 3 and 4 only at apical 1/7 or 1/6, anterior continuations of striae 3-5, whole of striae 1 and 2 and parascutellar striole not impressed. Granulation in marginal furrow discontinuous, interrupted briefly along second elytral quarter. Legs. Mesotibia evenly dilated apically. Male protarsomeres 1-3 expanded, 1 subelongate, nearly as long as wide, 2 and 3 wider than long; ventral adhesive setae present only at apical 1/3-1/4 surface of protarsomere 1 and entire surfaces of 2 and 3. Male basomesotarsus slightly flattened, with apical 2/3 provided with some short, stout, tooth-like setae ventrally; female basomesotarsus with finer setae ventrally. Female genitalia. Gonocoxite 2 subtriangular. Spermatheca narrow and long, nearly straight in basal 1/2, coiled in distal 1/2, undifferentiated; spermathecal gland connected near basal 1/6. Male genitalia. Median lobe of aedeagus in lateral view long, considerably bent ventrally, tapering distally, slightly broadened at apex (Fig. 34 View Figures 32–40 ), with relatively large basal bulb; lobe in dorsal view with apical lamella strongly bent to right, with right side quite concave and left side slightly and evenly convex (Fig. 24 View Figures 22–31 ); basal bulb forming edge dorsally with evident crest; sclerotized portion of endophallus visible in repose with three sclerites.

Notes.

Fauvel stated that this species has only four setae on the anterior edge of the labrum. However, the type specimen has a slightly malformed labrum with three setae on the right and two on the left. The median seta on the right is minute.