Cochlostoma (T.) dubium ( Wagner, 1897 )

Zallot, Enrico, Kamchev, Panche, Schilthuizen, Menno, Fehér, Zoltán, Mattia, Willy De & Gittenberger, Edmund, 2024, Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae), European Journal of Taxonomy 927, pp. 1-163 : 101-103

Cochlostoma (T.) dubium ( Wagner, 1897)

Figs 82 (green dots), 89–91

Pomatias (Auritus) nanus var. dubia Wagner, 1897: 55 , pl. 9 fig. 93a–b.

Types

Lectotype (here designated)

CROATIA • ♀; “ P. nanus W var. dubia Wagner orig. Metla bei Trnovac Croatien”; SMF160822 .

Paralectotypes

CROATIA • ♂; “ P. nanus Westerl. var. dubia A. J.Wagner, Metla bei Trnovac Croatien, Wagner det. et don.”; NHMW25358 /1 • 1 ♀ 4 ♂♂; “ P. nanus Westerl , var. dubia A. J.Wagner, Metla bei Trnovac Croatien ”; MIZPASW 7076 • ♂; “ P. nanus Westerl. var. dubia A. J.Wagner, Metla bei Trnovac Croatien , Wagner det. et don”; NHMW25358 /2 .

Other specimens

CROATIA • 1- Metla (topotypical); 44.645° N, 15.2459° E; 1900; unknown leg.; HNHM70300 • 2- Visočica peak 934; 44.4248° N, 15.331° E; 2008; Nádai leg.; HNHM97143 • 3- Baške Oštarije ; 44.5262° N, 15.1747° E; 1987; Kiss and L. Pintér leg.; HNHM 1987 /D022 • 4- Jakšina , Bijele Stijene; 45.2162° N, 14.9688° E; 1994; J. Steffek leg.; HNHM94017 • 5- Rossijev Kuk sp 1; 44.7643° N, 14.9882° E; 2007; L. Dányi and Á. Vári leg.; HNHM DL243 .

Type locality

CROATIA • Lika-Senj County, Trnovac, Mt Metla.

Description

SHELL. Closely spaced riblets on last part of protoconch. Teleoconch spotless, ribbed with not prominent ribs regularly spaced. Very developed and strong lip with indented columellar lobe covering umbilicus.

MEASUREMENTS. 8 ♀♀: whorls= 7.1–8.1, H = 7.2–8.4 mm, H/W= 2.28–2.67, roundness=0.11–0.14, ribs incl.= 53–56°, apert. incl.= 17–23°, ribs/mm 1 st wh.= 5–11, ribs/mm 4 th wh.=7–12.

FEMALE GENITAL ORGANS. Only badly preserved specimen from Visočica peak 934 was available for analysis of female genital morphology and it resulted as in C. (T.) nanum .

Remarks

We did not amplify the 16S of C. (T.) dubium . However, the shell and female morphology indicate that it is part of this clade. The shell similarity with C. (T.) nanum could be a convergent adaptation to an alpine environment. The male paralectotype NHMW25358/2 ( Fig. 90B ) differs substantially from the lectotype, not having the strong lip with an indented columellar lobe. It seems to be a specimen of C. (T.) elegans . Thus, NHMW25358/2 is not accepted as C. (T.) dubium .

Figures

Fig. 82.Distributionof samplesin Clade B: red= Cochlostoma (T.)braueri (Wagner, 1897);purple=C. (T.) stossichi (Hirc, 1881); blue=C. (T.) hallgassi sp. nov.; yellow =C. (T.) mariannae Nordsieck, 2011; green =C. (T.) dubium (Wagner, 1897); white = C. (T.) mostarense (Wagner, 1906). Black center for the samples with amplified 16S.

Fig. 90. Cochlostoma (T.) dubium (Wagner, 1897). A. Paralectotype (NHMW-25358/1). B. Supposed syntype, not conspecific with specimen A (NHMW-25358/2).