Coccopsilis recondita, Jaschhof & Jaschhof, 2020

Jaschhof, Mathias & Jaschhof, Catrin, 2020, The Porricondylini (Diptera: Cecidomyiidae: Porricondylinae) of Sweden revisited: descriptions of nineteen new species of ten genera, including a new genus, Zootaxa 4728 (2), pp. 151-182 : 155

Coccopsilis recondita sp. nov.

Figs 6–7

Diagnosis. Another typical Coccopsilis , C. recondita is characterized by male genitalic features, as follows ( Fig. 7 ). The gonostylus, which is twice as long as wide and slightly curved, has a large pectinate claw whose length equals the width (↓ 1). The edge of the claw is markedly rounded, for the spines in the center are longer than those at the margins. The apical edge of the tegmen is sinuous rather than evenly rounded due to the presence of a slightly protruding, horseshoe-shaped reinforcement (↓ 2). Coccopsilis recondita differs from C. obscura , a generally similar species, in the gonostylar claw being longer, narrower and more roundish; in the thicker aedeagal apodeme; and in the outline of the tegminal apex, which in C. obscura lacks the horseshoe-shaped reinforcement. Females and preimaginal stages of C. recondita are unknown.

Other male characters. Body size 1.3 mm. Head. Eye bridge 2–3 ommatidia long at vertex. Antenna longer than body, almost as long as wing. 12 flagellomeres; apical flagellomere long, composed of two bodies; circumfila on flagellomeres 1–11. Neck of fourth flagellomere 2.0–2.2 times as long as node ( Fig. 6 ). Palpus slightly shorter to slightly longer than head height, 4 setose, subcylindrical segments, apical segment longest of all. Thorax. Pronotum with 1–3 setae; anepisternum with 0–2 setae; anepimeron with 2–4 setae. Wing. Length / width ratio 3.0–3.3. Legs. Foreleg: femur slightly longer than tibia, tibia 0.9 times as long as T 2. Genitalia ( Fig. 7 ). Gonocoxal synsclerite: processes slightly bent, apices curved ventrally; dorsoposterior portions broad, angular, not appreciably prolonged posteriorly. Gonostylus with dent above the apical claw. Aedeagal apodeme about as long as parameres, moderately sclerotized with membranous apex.

Etymology. The Latin adjective recondita means hidden, an allusion to the fact that this species may easily be mistaken for C. obscura .

Type material. Holotype. Male , Sweden, Småland , Nybro , Bäckebo, Grytsjön Nature Reserve, old-growth mixed hemiboreal forest, 12 August–6 October 2014, Malaise trap, M. & C. Jaschhof (spn. CEC 2544 in NHRS).

Paratypes. 1 male, same locality as the holotype, 2–12 July 2005, MT, SMTP (trap 1000, collecting event 1323) (spn. CEC 2546 in NHRS) ; 1 male, same locality but hay meadow at forest edge, 13–24 July 2005, MT, SMTP (trap 1001, collecting event 1355) (spn. CEC 2547 in NHRS) ; 1 male, same data as the holotype but 17 July–11 August 2014 (spn. CEC 2543 in SDEI) ; 1 male, same data but 30 September–30 November 2015 (spn. CEC 2545 in SDEI) .

Other material. Sweden: 1 male, Öland , Mörbylånga, Ottenby lund (military radar station), old-growth mixed broadleaf forest, 20 August–17 September 2004, MT, SMTP (trap 21, collecting event 995) (spn. CEC 2542 in NHRS) .

Figures

FIGURES 6–9. Male morphology of Coccopsilis recondita (6–7) and Coccopsilis scalpta (8–9), holotypes. 6: Fourth flagellomere, lateral. 7: Genitalia, ventral. 8: Genitalia, ventral. 9: Fourth flagellomere, lateral. Scales 0.05 mm. Arrows refer to characters described in the diagnoses.