Clistoabdominalis crassus ( Bezzi, 1926 )

FÖLDVÁRI, MIHÁLY, 2013, Taxonomic revision of the Afrotropical species of the tribe Eudorylini (Diptera, Pipunculidae), Zootaxa 3656 (1), pp. 1-121 : 18-20

Clistoabdominalis crassus ( Bezzi, 1926)

( Figs 5F–K , 40J )

Pipunculus crassus Bezzi in Bezzi and Lamb 1926: 550 — comb. nov.

Diagnosis: Variability of male genitalia is unknown as the species is known from only a single male and female. Surstyli with protuberances at base, SES not developed; gonopods not clear, hypandrium deflected in ventral view and a sclerite seems to be attached to it.

Type material: Mauritius: 1♂, LT, Rodriguez I., viii–ix. 1918, HP Tommasset and HJ Snell; [separate label:] Rodriguez I., Pres by Dr H Scott, BM 1926-190; 1♀, PLT, same data as LT ( BMNH) .

Male

Head. Third antennal segment acuminate; brown, tip whitish. Face silvery pollinose. Frons, upper part shining black, lower part silvery pollinose; eyes touching for distance equal to 2 times ocellar triangle. Occiput brownish pollinose.

Thorax. Humeri yellow-brown. Mesonotum (viewed obliquely from front) and scutellum subshining (but surface is greasy); mesonotum brownish (slightly pollinose) from the side. Scutellum with 2–3 very weakly developed hairs. Dorsocentral hairs weakly developed. Halter yellow, stem darker.

Legs. Trochanters and femora brown, f1,2 posteriorly silvery pollinose; knees, tibiae and tarsal segments yellow, last segment brown. Ventroapical row of spines missing on femora. Subapical (distal) spines on first four tibiae present. No anteromedial hairs on 3rd tibia. Hind trochanter with a row of 10–12 dark spines (as a comb) on ventral side. Pulvilli shorter than last tarsal segment.

Wing. Cross-vein R-M at 2/5 of discal cell. 4/5 of pterostigma coloured. Hairs on tegula absent.

Abdomen. Tergites black with brownish pollinosity (without any grey colour). No hairs on abdomen, no lateral spines on first tergite (may be broken). Postabdomen in dorsal view: EP visible; T5 0.8 times as long as ST8. Genitalia without dissection: no m.a., SS yellow.

Genitalia. Surstyli with protuberances at base; ST8 large, but damaged ( Fig. 5G ). SES not developed; gonopods not clear, the pointed structure in the middle must be the pg; hypandrium deflected in ventral view and a sclerite seems to be attached to it ( Fig. 5I ); phallus uncertain, may be lost in LT; ejaculatory apodeme mushroom shaped; sperm pump funnel-like ( Fig. 5K ).

Female

As male except for the following characters. Frons, eyes separated; silver-grey pollinose on lower 1/3 and along the margins up to the middle, otherwise shining black. Subapical (distal) spines on first four tibiae very distinct. Pulvilli and claws about 1.5 times as long as last tarsal segment. Female ovipositor ( Fig. 40J ) with thin, slightly upcurved piercer.

Remarks –The male specimen is hereby designated lectotype of C. crassus to fix the name and ensure a universal and consistent interpretation of the taxon.

Distribution— Mauritius.

Figures

FIGURE 5. Male genitalia of Clistoabdominalis nitidifrons (A–E) and Cli. crassus (F–K).—A: outer surstylus with phallic guide, lateral view, B: dorsal view, C: inner surstylus, lateral view, D: ventral view with phallic guide, sperm pump and ejaculatory apodeme (without ST8), E: surstyli, dorsal view, F: outer surstylus with phallic guide, lateral view, G: dorsal view, H: inner surstylus, lateral view, I: ventral view with phallic guide, (without ST8), J: surstyli, dorsal view, K: sperm pump with ejaculatory apodeme. Scale bars: 0.1 mm

FIGURE 40. Female ovipositors.—A–B: Eudorylas aethiopicus, C–D: Eudorylas bredoi, E: Clistoabdominalis lomholdti, F: Clistoabdominalis nitidifrons, G: Eudorylas falcatus, H–I: Eudorylas flexus, J: Clistoabdominalis crassus, K: Eudorylas setiformis, L: Clistoabdominalis lomholdti, M: Dasydorylas sordidatus, N–O: Eudorylas mikenensis, P–Q: Eudorylas decorus, R–S: Eudorylas galeatus, T: Eudorylas sinuosus, U: Eudorylas wittei, V–W: Dasydorylas evanidus. Scale bar: 0.5 mm