Claraeola nigripennis (Hardy, 1949)

FÖLDVÁRI, MIHÁLY, 2013, Taxonomic revision of the Afrotropical species of the tribe Eudorylini (Diptera, Pipunculidae), Zootaxa 3656 (1), pp. 1-121 : 14-15

Claraeola nigripennis (Hardy, 1949)

( Figs 4A–F )

Congomyia nigripennis Hardy 1949 b: 7 .

Diagnosis: Large species, wings dark brown. EP small, m.a. wide, SES well developed hairy, PG pointed, with minute hairs just below apex, phallus trifid with apical projection covered by scale-like setae.

Type material: Democratic Republic of Congo: 1♂, HT, Congo-belge, Eala - III. - 1936, J. Ghesquière, R. Mus. Hist. Nat. Belg. I.G.10.482 ( ISNB).

Other material examined: Cameroon: 1♂, Mission Cameroun, CNRS, RCP 318, Octobre Novembre 1975, Bafut Nguemba (Vallee de la Haute Nguemba), Province du Nord-Ouest, 3.XI.1975, “ Eudorylas sp. det. J. Skevington / 98” ( MNHN) . Democratic Republic of Congo: 1♂, Kasai: Terr. de Dekese Itunda, XII. 59, F.J. François, R. I. Sc. N. B., I.G. 24.452 ( ISNB) . Uganda: 1♂, Kilembe., 4,500 ft., F. W. Edwards.; Ruwenzori Range., xii.1934 – i.1935., B.M.E. Afr. Exp., B.M. 1935-203 ( BMNH).

Male

Head. Third antennal segment obtuse, dark brown, pedicel with strong black bristles ventrally and dorsally. Face and frons brownish pollinose (also in front of ocellar triangle); eyes touching for distance equal to 2.5 times ocellar triangle. Occiput, lower half silvery pollinose along the edges, otherwise and upper half subshining black.

Thorax. Humeri dark brown. Mesonotum (viewed obliquely from front) brownish pollinose, also brownish from the side, notopleural depression with numerous dark hairs. Scutellum brown, with few pale hairs. Dorsocentral hairs well developed. Halter dark brown.

Legs. Trochanters, femora, knees and tibiae dark brown. Tarsal segments yellow brown, last segment darker (f1,2 silvery pollinose pv). Tarsi of the 3rd leg flattened. Ventroapical row of 5–8 short, black spines on all femora (small compared to the size of the specimen), all femora with a row of hairs (posteriorly) at least as long as width of femora at base. Subapical (distal) spines on first four tibiae missing. The anteromedial hairs slightly erected along distal half of the 3rd tibia. Hind trochanter with 6–10 short, dark spines on ventral side. Pulvilli distinctly shorter than last tarsal segment.

Wing. Dark brown, fumose (may be paler, shorter). Fourth costal section 0.5–0.8 times as long as third costal section. Cross-vein R-M at 2/5 of discal cell. Pterostigma fully coloured. Hairs on tegula uncertain.

Abdomen (previously dissected). (based on specimen from Kasai) Viewed obliquely from front tergites 2–4 subshining-shining black, T1 silvery pollinose dorsally. Hairs dispersed, short and weakly developed in general. Laterally longer dark hairs (20–25) on first and second tergite as long as width of t3 at base.

Genitalia. Surstyli rectangular, IS larger (up to twice as large as OS); epandrium small compared to ST8, which bears a wide m.a., as large as ¾ of width of ST8 (may be smaller) ( Fig. 4B ). SES well developed, with many hairs on both sides ( Fig. 4F ); gonopods asymmetrical, IG small, round, OG larger, more rectangular; hypandrium broad; phallic guide pointed apically, with minute hairs below apex (also on dorsal side) ( Fig. 4A ); phallus trifid with apical projection covered by scale-like setae; ejaculatory apodeme (based on specimen from Cameroon) linear, flat; sperm pump small, round, with two patches of hairs ( Fig. 4D ).

Female—Unknown.

Distribution— Cameroon, Democratic Republic of Congo, Uganda.

Figures

FIGURE 4. Male genitalia of Claraeola nigripennis (A–F) and Cla. stuckenbergi sp. nov. (G–K).—A: outer surstylus with phallic guide, lateral view, B: dorsal view, C: inner surstylus, lateral view, D: sperm pump with ejaculatory apodeme, E: surstyli, dorsal view, F: ventral view with phallic guide, phallus (without ST8), G: outer surstylus with phallic guide and outer gonopod, lateral view, H: dorsal view, I: inner surstylus, lateral view, J: surstyli, dorsal view, K: ventral view with phallic guide, phallus (without ST8). Scale bars: 0.1 mm