Cladiopsocus tinigua, Obando & García Aldrete & Carrejo, 2018

Obando, Ranulfo González, García Aldrete, Alfonso N. & Carrejo, Nancy, 2018, New species and records of Cladiopsocus Roesler (Psocodea: ‘ Psocoptera’: Psocomorpha: Cladiopsocidae) from Colombia, Zootaxa 4378 (2), pp. 191-223 : 213

Cladiopsocus tinigua n. sp. Male

( Figs 83–89 )

Diagnosis. Belonging in species group “1”, subgroup 2 in the classification of Eertmoed (1986). The wings, phallosome, epiproct and paraprocts relate it to C. amacayacuensis González et al. , but differs from it in having the paraproctal prong well developed and tapered distally ( Fig. 89 ). Epiproct posteriorly emarginated, postero-lateral corners widely rounded ( Fig. 88 ).

Color. Head ( Fig. 85 ) brown, vertex, labrum and mandibles brown, maxillae pale brown, labium cream. Compound eyes dark brown, ocelli hyaline in a dark brown area. Antennae and maxillary palps pale brown, flagellomeres cream apically. Pro-, meso- and metanotum pale brown, meso and metascutellum dark brown, thoracic pleurae and legs pale brown, tibiae and tarsi darker. Forewing membrane hyaline, with small, slightly smoked areas ( Fig. 83 ); veins brown, with pale brown spots along the veins; wing margin with brown spots, two on areola postica. Hindwings hyaline ( Fig. 84 ). Abdomen pale brown to cream, incomplete abdominal segmental rings pale brown, clunium and hypandrium brown, clunium with narrow anterior edge dark brown; phallosome and epiproct pale brown; paraprocts brown, paraproctal prong darker distally; laminar projection between the prong and tubercle dark brown.

Morphology. As in diagnosis plus the following: head vertex slightly emarginate, almost at the same level of the upper margin of the compound eyes ( Fig. 85 ); these large, with interommatidial setae. D is 0.55 the length of the head in front view. Forewings ( Fig. 83 ); membrane pterostigma with setae, without spur veins. Hypandrium triangular, broad, anteriorly concave, with medium sized setae and macrosetae ( Fig. 86 ). Phallosome short, piriform ( Fig. 87 ); endophallic lobes broad, rounded apically, separate from each other along the midline; external parameres rounded, membranous, bearing sensorial cones; internal parameres broad, curved, fused in the middle, with blunt apices directed posteriorly. Epiproct semioval, wider than long, emarginate posteriorly ( Fig. 88 ), with three macrosetae mesally and two near each postero-lateral corner, posterior setal field as illustrated. Paraprocts ( Fig. 89 ) with medium sized setae and macrosetae on long, stout mesal process as illustrated; a large, sclerotized, rounded protuberance between the mesal process and the prong, this wide proximally, short, acuminate; sensory fields with 19 trichobothria on basal rosettes ( Fig. 89 ).

Measurements. FW: 2900, HW: 2125, F: 625, T: 1125, t1: 425, t2: 52, t3: 62, ctt1: 22, f1: 525, f2: 387, f3: 325, f4: 275, f5: 170, f6: 162, f7: 137, f8: 125, f9: 107, f10: 100, f11: 95, Mx4: 165, IO: 205, D: 330, d: 230, IO/d: 0.89, PO: 0.70.

Specimens studied. Holotype male. COLOMBIA. Caquetá. Belén de Los Andaquíes, Resguardo Indígena La Esperanza, 1°36’19.18’’N : 75°56’12.46’’W, 844 m. J. Panche. Led light trap in forest canopy. MUSENUV slide code 2897. Paratypes: 1 male, same data as the holotype . 4 males, Caquetá. San Vicente del Caguán, Laureles, Resguardo Indígena Altamira, 2°27’50.14’’N : 74°55’02.06’’W, 917 m. 27.IV.2017. Led light trap in forest canopy. J. Panche. 5 males, same locality and collector. 26–29.III.2017 .

Etymology. The specific epithet honors the Tinigua ethnic group, now extinct, that inhabited the basins of the rivers Yarí, Caguán and Guayabero, in the department of Caquetá.

Figures

FIGURES 83–89. Cladiopsocus tinigua n. sp. Male. 83. Forewing. 84. Hindwing. 85. Front view of head. 86. Hypandrium. 87. Phallosome. 88.Epiproct.89. Paraproct.Scales in mm.